Harris: Male antennule of porcellidiid copepods 
113 
Key to the species of Ravania 
1 Female caudal furca “dolioid” (barrel-shaped), rami not rhom¬ 
boid. (Plate 1E,F, p. 163) . Ravania doliocauda sp. nov. 
- Female caudal furca pointed, rami rhomboid with T4 at rounded 
apex .2 
2 Anterior half of female genital double-somite without border 
setules. Male P2 endopod with two terminal setae. Short rows of 
setules at base of each terminal seta on male P5. (Plate 1 A,B). Ravania wellsi sp. nov. 
- Anterior half of female genital double-somite with border 
setules. Male P2 endopod with three terminal setae. No setules 
at base of each terminal seta on male P5. 
. Ravania ravanae (Thompson & Scott, 1903) comb. nov. 
Ravania wellsi sp. nov. 
Figs 1-4, Plate 1A 
Type material. Holotype adult male, length 0.51 mm, 
P81198; allotype adult female, length 0.58 mm, P81199; 
paratype specimens, 5 3 SS, 3 juveniles, P81200, 
deposited at AM, Sydney. Additional paratypes deposited at 
NHM, London. All collected from Eucheuma denticulatum , 
sublittoral fringe, Point Vernon, Hervey Bay, Queensland 
(25°15'S 152°47'E), V. A. Harris, 1998. 
Diagnosis. Male antennule with short pointed spur or 
thom-like denticle at base of 8 seta on segment 3; anterior 
half of female genital double-somite without border 
setules, posterior half with border setules, no seta in lateral 
indentation between anterior and posterior lobes, half caudal 
ramus contained in arch of genital double-somite; female 
caudal ramus rhomboid, terminal seta T2 almost as long as 
y seta; male P2 with only two terminal plumose setae on 
endopod segment 3; female P4 with two spinous setae on 
endopod; female P5 exopod with one dorsal and two apical 
setae; male P5 with short rows of ventral setules at base of 
each terminal seta. 
Biometric data. Females (N = 29): maximum length (L m ax) 
mean 0.57 mm, range 0.54-0.61 mm, body length (L urs ) mean 
0.52 mm, range 0.49-0.56 mm; cephalosome width (W) mean 
0.36 mm, range 0.34—0.44 mm; rostrum width 0.08 mm; width 
of genital double-somite 0.23 mm, length 0.14 mm, arch 0.08 
mm; caudal ramus 0.115 mm, width 0.042 mm. 
Ratios: L U rsAV 1.44, W/R 4.5; genital double-somite w/1 
1.65, arch 57% of length; caudal ramus 22% of L urs, ramus 
1/w 2.7, Hicks’ index for (3 68%. 
Males (N = 23): length (L m ax) mean 0.51 mm, range 
0.49-0.54 mm, body length (L U rs) mean 0.48 mm, range 
0.46-0.51 mm; cephalosome width 0.35 mm. antennule 
(N = 3, fully extended 0.125 mm; spermatophore 0.18 x 
0.065 mm. 
Ratios: L U rsAV E4; antennule 24.5% of Lurs, antennule 
segment 2 31%, segment 3+4 35%, dactylus 21% of 
antennule length; spermatophore 37% of body length L U rs.. 
Description. Adult females (Fig. 1A; Plate 1A, p. 163): 
colouration, pale red with central cephalosome and 
metasome segments 1 and 2 deep red, metasome segments 
3 and 4 pale red, genital double-somite and caudal rami 
deep red. Some animals are pale red or colourless. Eye spot 
dark brown. Anterior of cephalosome broadly rounded, 
rostrum prominent, narrow with hyaline anterior border. 
Dorsal surface ornamented with pits (4 pm). Hyaline border 
8 pm wide, without striations. Sensillum on epipleural lobe 
of second metasome segment conspicuous, longer than 
other species. Labrum without central patch of setules or 
ridge plates. Genital double-somite (Fig. IE) with slight 
indentation that marks the boundary between anterior and 
posterior lobes, anterior lobe short, convex, with thickened 
edge but no setules, posterior lobe triangular, pointed 
posteriorly with border setules. Arch of genital double-somite 
deep, encloses about half length of caudal ramus, dorsal 
surface with pits. Genital opening as in Fig. IF. Caudal 
rami (Fig. ID) rhomboid, length about 2 Yz times width, 
sides almost parallel, oblique posterior edge slightly convex, 
apex broadly rounded with pinnate T4. Dorsal surface with 
prominent network of ridges, |3 seta A way down ramus, 
y seta about Yz way along oblique edge. Medial edge with 
border setules up to the level of (3 seta, lateral edge slightly 
convex with border setules distally, T1 unipinnate at lateral 
corner of oblique edge, T2 plain about as long as y seta, 
T3 absent, posterior border with fine setules from T1 to 
T4. Structure and setation of mouthparts and ambulatory 
limbs typical of family. Antenna (Fig. 2A) exopod with 
five plumulose setae and one finely serrulate spinous seta, 
endopod segment 2 with two lateral setae, geniculate setae 
with plain end segment, claw fine comb-like. Mandible (Fig. 
2D, E, F) without setules on anterior lobe of palp. Maxillule 
(Fig. 2B), endopod with six setae, exopod with one bulbous 
seta. Maxilla (Fig. 2H) and maxilliped (Fig. 2G) typical. 
No area of denticulate setules on PI endopod (Fig. 2C). 
Serrulate spinous seta on segment 3 of P2 endopod about 
% length of endopod (Fig. 3C). Serrulate spinous seta on 
segment 2 of P3 endopod (Fig. 3A) shorter than endopod 
(0.8:1), large serrate spinous seta on segment 3 longer than 
endopod (1.3:1). Internal seta on segment 2 and segment 
3 of P4 endopod strong spinous, other setae plumose (Fig. 
3F). Dorsal surface of P5 exopod with pits, internal seta on 
baseoendopod serrulate, about % length of exopod, exopod 
lanceolate with one dorsal seta and two apical setae (Fig. 
3E). P5s reach beyond genital double-somite to extremity 
of caudal rami, but do not touch posteriorly. Females carry 
six large eggs. 
Adult males { Fig. IB; Plate lB,p. 163). Colouration similar 
to female, but posterior region of most specimens is pale red or 
colourless. Anterior of cephalosome rounded (not truncated), 
rostrum projects anteriorly, in ventral view rostrum appears 
