Harris: Male antennule of porcellidiid copepods 
117 
characteristic of genus. Seta on segment 1 plumose, segment 3 
with short, pointed spine-like denticle close to anterior lobe, 5 
seta very long (A length of antennule), projects anteriorly (Fig. 
21), two tooth-like coupling denticles on segment 4, aesthetasc 
extremely long (almost as long as fully extended antennule 
(Fig. 21), dactylus long, cylindrical. Endopod of P2 with two 
plumose terminal setae (Fig. 3B). No spinous setae on P4 
endopod, all setae plumose. Dorsal surface of P5 exopod with 
pits, ventral setules at base of each terminal seta (Fig. 3D). 
Etymology. The species has been named in recognition 
of Dr J. B. J. Wells’ contribution to the taxonomy of the 
Harpacticoida. 
Distribution. Type series collected from Eucheuma 
denticulatum, Point Vernon, Hervey Bay, PV4. 7/98,41 $ $ 
(21 with eggs), 26 SS, 14 juveniles. Also found on Zonaria 
sp., PV8, 8/00, 12 29, 14 $$ at the same location. The 
species has been recorded at Arrawarra headland, northern 
NSW, on Caulerpa vesiculosus and Dilophus sp. V. A. Harris, 
1982, 2000. 
Ravania doliocauda sp. nov. 
Figs 4-8, Plate IE 
Type material. Holotype adult male, length 0.66 mm, 
P81195; allotype adult female, length 0.75 mm, P81196; 
paratype specimens, 10 6 SS-, P81197, deposited 
at AM, Sydney. Additional paratypes deposited at NHM, 
London. All collected from Zostera capricornia at Pulgul 
Creek mud flats, Urangan, Hervey Bay, Queensland (25° 17'S 
152°52'E), V. A. Harris, 30 Nov. 2002. 
Diagnosis. Large denticle with serrated crown at base of 8 
seta on segment 3 of male antennule; female genital double¬ 
somite narrow (< A cephalosome width), no indication 
of division into anterior and posterior lobes, narrows to a 
point posteriorly, border setules absent except for a few at 
posterior apex, caudal rami excluded from posterior arch of 
genital double-somite; caudal furca dolioid (barrel-shaped 
in outline), rami long (1/w > 2), taper posteriorly, lateral and 
posterior edge convex, T1 lateral (A way down ramus), y seta 
recessed at posterolateral comer; medial patch of setules on 
labrum; male P2 endopod with three terminal setae. 
Biometric data. Females (N = 32): maximum length (L m ax) 
mean 0.75 mm, range 0.72-0.78 mm, body length (L U rs) 
mean 0.65 mm, range 0.62-0.69 mm; cephalosome width 
(W) mean 0.45 mm; rostrum width (R) 0.10 mm; genital 
double-somite length 0.135 mm, width 0.23 mm, arch 0.03 
mm; caudal ramus length 0.13 mm, maximum width 0.06 
mm (A down ramus). 
Ratios: L U r S /W 1.44; W/R 4.5-4.75; genital double-somite 
width 50% of body width, arch 22% of length; caudal ramus 
20% of L U rs, ramus 1/w 2.16, terminal seta T1 located 54% 
down lateral edge of ramus, Hicks’ index for a 85%, for (3 70%. 
Males (N = 20): maximum length (L m ax) mean 0.66 mm, 
range 0.62-0.67 mm; cephalosome width (W) mean 0.44 
mm; rostrum length 0.055 mm (ventral); caudal ramus length 
0.05 mm, width 0.05 mm; antennule length fully extended 
0.165 mm; spermatophore 0.22 x 0.09 mm. 
Ratios: L m ax/W 1.5; caudal ramus 7.5% of Lmax, ramus 
1/w 1.0; antennule 20% of L max , antennule segment 2 24%, 
segment 3+4 36%, dactylus 24% of antennule length; 
spermatophore 33% of body length Lmax. 
Description. Adult females (Fig. 5A; Plate IE, p. 163): 
colourless or very pale yellow (but see remarks below), 
outline of body oval, cephalosome semicircular with 
prominent rostrum, dorsal pits inconspicuous, very few 
dorsal sensilla mostly towards edge of cephalosome. Hyaline 
border clear, 7 pm wide (Fig. 5F). Labrum with central patch 
of very short setules (Fig. 6J). Genital double-somite (Fig. 
5B, C) narrow, no epipleural expansion or division into 
anterior and posterior lobes, posterior half narrow (equals 
width of caudal rami), very fine filiform setules towards 
acutely pointed posterior, dorsal surface with transverse 
rows of shallow pits, no cleft, notch or scar to indicate 
boundary between anterior and posterior regions. Caudal 
arch shallow, encloses most of the anal segment, but caudal 
rami are excluded from arch. Female genital opening (Fig. 
5D, E). Caudal furca dolioid (barrel-shaped in outline) (Fig. 
5B). Each ramus narrows posteriorly, maximum width at 
level of (3 seta, lateral edge convex with border setules, 
medial edge without setules, posterior edge slightly convex, 
dorsal surface with pits and reticulate marking posteriorly 
(Fig. 61).Terminal seta T1 halfway down lateral edge, y seta 
recessed, T4 set in from medial comer. T3 absent. Antennule 
with pinnate seta on segment 1. Structure and setation of 
mouthparts and ambulatory limbs typical of family. Basis 
of antenna with row of small triangular setules (Fig. 6D), 
segment 2 of endopod with two lateral setae, geniculate 
setae with plain end segment, claw with fine comb-like 
edge. Mandible (Fig. 6H), maxillule (Fig. 6A), maxilla 
(Fig. 6C) andmaxilliped(Fig. 6B). Segment 1 of PI exopod 
(Fig. 6G) with crescent of fine setules parallel to edge, no 
area of denticulate setules on endopod. Spinous seta on P2 
endopod segment 3 more than A endopod length (0.6:1) 
(Fig. 1C). Serrate spinous seta on segment 2 of P3 endopod 
almost as long as endopod (Fig. 7B), large serrate spinous 
seta on segment 3 longer than endopod (1.4:1). P4 endopod 
with internal spinous seta on segment 2 and first (internal) 
spinous seta on segment 3 plain (Fig. 7E). Baseoendopod 
of P5 with long ventral seta (A length of exopod), exopod 
broad (1/w = 1.67), ovate with acute apex (Fig. 6E), strong 
ventral falciform ridge, long border setules, one subterminal 
dorsal seta and two apical setae (Fig. 6F), P5s extend back 3 A 
length of caudal rami and compensate for the narrow genital 
double-somite by forming most of the roof to the brood 
chamber. Females carry 10 eggs per brood. 
Adult males (Fig. 8A; Plate IF, p. 163). Colourless or very 
pale yellow. Outline of body oval, anterior of cephalosome 
semicircular (not truncated), rostrum keeled, oval in ventral 
view (Fig. 8C). Dorsal pits, sensilla and hyaline border 
as for female. Caudal ramus (Fig. 8E) quadrate, setation 
similar to female T1 lateral, T3 absent, T4 at medial corner. 
Antennule (Fig. 7F) with pinnate seta on segment 1, 8 seta 
on segment 3 long (equal to segment 3+4+ dactylus in 
length), points forward, coupling denticle on segment 3 large 
with serrated crown, two tooth-like denticles on segment 4 
(Fig. 8F), dactylus As length of segment 3+4, apex pointed 
(Fig. 8G). Other limbs as for female except P2 (Fig. 7D) 
has three terminal setae on endopod (two plumose, one 
serrulate spinous). P5 acutely trapezoidal, first (lateral) seta 
of different shape to terminal setae (Fig. 8D), first row of 
ventral setules about 16, row of four or five setules at base 
