Harris: Synopsis of genera and species 
183 
Genus Clunia gen. nov. 
Type species. Clunia cocosensis sp. nov. 
Diagnosis. Maxillule endopod with only one seta;female 
cephalosome not truncated; hyaline border with sensilla at 
lateral edge of cephalosome; no dorsal organs with cuticular 
honeycomb; no ridge plates on labrum; no lateral striations to 
anterior lobe of female genital double-somite; female caudal 
ramus pentagonal, median corner 90°, posterior border 
straight, T3 absent; coxae of maxillipeds touch in midline; 
no ventral expansion to female P5 exopod. 
Species composition. Clunia cocosensis sp. nov., is the only 
species currently known. Recorded from the Indian Ocean. 
Etymology. The genus is named Clunia after John Clunies- 
Ross who settled on the Cocos (Keeling) group of islands 
in 1827. 
Remarks. The shape of the caudal ramus and absence of T3 
resemble Geddesia and Mucrorostrum , but the presence of 
only one seta on the maxillule endopod (in contrast to six on 
the former and two on the latter) and the endite formula are 
unique features that justify erection of the genus Clunia. No 
male specimen is available for male characters that would 
confirm this decision. 
Clunia cocosensis sp. nov. 
Figs 10-11 
Type material. Holotype adult female, length 1.06 mm, 
P81219; paratype specimens (adult female and dissected 
female on slide) P81220, deposited at AM, Sydney. All 
collected from Cocos (Keeling) Islands, Indian Ocean, F. 
H. Talbot, 1979. 
Diagnosis. Female cephalosome hemi-elliptical; dorsal 
cuticle almost devoid of pits; anterior lobe of mandibular 
palp long, without ventral setules, molar process modified as 
a scraper; single seta on maxillule exopod long, not bulbous, 
endopod with one seta, endite formula 2-3-1; first dorsal 
seta on P5 exopod close to lateral seta, apical seta pinnate. 
Biometric data. Females (N = 3): maximum length (L max ) 
I . 03,1.06,1.09 mm, body length (L U rs) 0.94,0.97,0.99 mm; 
cephalosome width 0.65 mm; rostrum width 0.137 mm; 
genital double-somite 0.35 mm wide, 0.24 mm long: caudal 
ramus 0.12 mm long, 0.08 mm wide. 
Ratios: L U rsAV 1.5, LmaxAV 1 .6, W/R4.75; genital double¬ 
somite w/1 1.46; caudal ramus 12.4% of L U rs, caudal ramus 
1/w 1.5, Hicks’ index for a 80%, for |3 68%. 
Description. Adult female (P81219, Fig. 10A). Natural 
colour unknown, specimens have light brown oil (?) droplets 
in body cavity. Rostrum not prominent, no lens in rostrum, 
dorsal pits small (3 pm) near edge of cephalosome, metasome 
segments and P5. Hyaline border clear, 12 pm wide. Genital 
double-somite (Fig. 11D) not broad, pointed posteriorly, 
lateral edge almost straight with no trace of division into 
anterior and posterior lobes, border setules absent except 
at apex, posterior arch less than % length of genital double¬ 
somite. Caudal ramus (Fig. 10G) pentagonal, maximum 
width about % down ramus, posterior border straight, 90° 
to medial edge. A diagonal ridge runs from proximal medial 
corner to insertion of T2 seta. Terminal setae Tl, T2, T4 
pinnate, Tl and y recessed on beveled edge, T4 at medial 
corner, T3 absent. Terminal fringe of setules between T2 and 
T4 % maximum width of ramus. Structure and setation of 
mouthparts and ambulatory limbs typical of family. Exopod 
of antenna with five plumulose setae and one spinous seta, 
endopod segment 2 with three lateral setae, first terminal seta 
short, geniculate setae with plain end section, claw comb-like 
(Fig. 10E). Cutting edge of mandible molar process modified 
into trowel-like scraping organ without anterior seta or lappet 
(Figs 10B, C). Precoxa of maxillule elongate, endites bear 
2-3-1 setae respectively, single plain seta on endopod and 
exopod (Fig. 10D). Maxilla (Fig. 1 IF) claw on endopod not 
broad or serrate, maxilliped (Fig. 10F). Conspicuous crescent 
of setules on segment 1 of PI exopod (Fig. 11E), endopod 
with small but conspicuous triangle of setules at lateral end 
of fimbriate crescent. Spinous setules on external edge of 
P2, P3 and P4 exopod segments 1 and 2 appear to lie in a 
double row of about 9 + 9 setules (Figs 11 A, C, G), segment 
1 has a proximal row of nine fine setules and segment 3 
has five or six setules. Serrated spinous seta on segment 2 
of P3 shorter than endopod, large serrated spinous seta on 
segment 3 strong (Fig. 11C) longer than endopod (1.4:1). 
Spinous setae on P4 endopod segments 2 and 3 plain (Fig. 
11G). P5 exopod lanceolate, not truncated posteriorly, two 
dorsal setae (first located very close to lateral seta—a position 
not known elsewhere in the Porcellidiidae) and one pinnate 
apical seta (Fig. 11B), border setules strong, P5s reach just 
beyond genital double-somite. One of the specimens has 
four large eggs in the brood chamber. 
Adult male (no specimen available). 
Etymology. Named from the Cocos Islands where the 
species was collected. 
Remarks. The absence of a male animal renders the above 
description incomplete. The position of the first dorsal seta 
close to the lateral seta on the female P5 is unusual and has 
not been observed on any other member of the family. 
Distribution. The four female specimens were collected at 
a depth of 4 m from dead coral encrusted with algae in a 
channel at the reef crest between Direction Island and Prison 
Island, Cocos (Keeling) Islands, Indian Ocean, F. H. Talbot, 
8 Oct. 1979. 
