192 
the mesenterial filaments. The oral cone is inclined distally so that a vertical section attains 
the actinopharynx obliquely. The ectoderm is very much folded; so its thickness varies from 
35 , x — 90 p. There is no connection between the number of folds and the mesenteries; two 
folds only, one on each sagittal end, are constant. The nervous layer is well developed; the 
muscular layer contains a few longitudinal fibres only. There are a large number of deeply 
staining granular glandcells, which very often consist of a peripheric part and a part in the 
lower epithelial half, both parts connected by a filiform stalk (PI. II, fig. 7). There are numerous 
hyaline glandcells at every depth of the ectoderm. The ectodermal surface stains deeply and is 
vertically striated, sharply limited from the deeper epithelium (PI. II, fig. 7). This surface-layer is 
probably formed by bundles of ciliae. There are no nematocysts. The mesogloea, a few p. 
thick, is somewhat thickened at the places of attachment of the mesenteries. It is a homogeneous 
layer. The entoderm (55 u) contains only a few deeply staining glandcells, not granular and 
less deeply stained than in the ectoderm. The upper part of the actinopharynx is entirely free 
from pigment, but in the lower part pigmentation appears, increasing in quantity towards the 
free actinopharyngeal border. This is very obvious in PI. V, fig. 7 where the section is oblique; in 
this figure the sagittal part of the actinopharyngeal ectoderm is free from pigment; its quantity 
increases towards the middle of the figure where a lower part of the actinopharynx is attained. 
Mesenteries (PI. II, figs. 8 and ii; PI. V, figs. 3, 4, 5, 7 and 24). In the number of 
mesenteries Eiicirripathes contorta differs from all other described species of this and other 
genera except the Eucintipaihes- species, which formerly was called Leiopathes. There are also 
typical differences in ovaria and mesenterial filaments, which render this species rather curious. 
On a horizontal section ot the upper part of the actinopharynx 
(PI. V, fig. 5) there are the normal number of the mesenteries visible, 
like in other species. But halfway down the oral cone an extra pair 
of secondary mesenteries appears (PI. V, fig. 7), situated between the 
normal pair of secondary mesenteries and the primary sagittal mesen¬ 
teries, while in Leiopathes this 6 th pair of mesenteries is found by 
Brook and van Beneden between the normal secondary ones and 
the primary transversal mesenteries. While all the other pairs of 
mesenteries reach the actinopharyngeal wall, this is not the case with 
the sixth pair, which is attached to the bodywall only (!). This sixth pair 
descends to the same depth as the other secondary mesenteries, even 
deeper than the primary sagittal mesenteries. — For the greater part 
the structure of all the mesenteries is the same as elsewhere. But in 
regularly distributed places the mesogloea bears lamellae of the same 
type as the lamellae which support the musclefibres in Actinia. Here 
Fig. 256. Eucirripath.es co/itorta v. a ] so q iese lamellae support a slightly developed system of longitudinal 
Pesch. System of the longitudinal mus¬ 
clefibres on the mesenteries (for the text musclefibres. These lamellae are clearly visible in PL V, figs. 3 and 
24; they are arranged unilaterally on the mesenteries according to 
the following system (PI. V, fig. 24; textfig. 256) in all the polyps: 
on the anterior pair of primary sagittal mesenteries the longitudinal musclefibres are found on 
cf. the list of abbreviations, before the 
plates). 
