16 
The Bulletin 
IV. Investigations Regarding the Inheritance of Types of Flowers 
in Vitis Rotundifolia 
With, the discovery of the variety, Hope, we find that Vitis rotundi¬ 
folia produces three distinct types of flowers instead of two. The im¬ 
portant question now arises as to how and in what manner these three 
types of flowers are transmitted to the progeny of the first generation. 
Hedrick and Anthony (3), from tabulations of work done with many 
seedlings of Euvitis, report the following ratios: 
Upright stamens X upright stamens=4.3 upright: 1 reflexed 
Reflexed stamens X reflexed stamens=1.2 upright: 1 reflexed 
Reflexed stamens X upright stamens=l. upright: 1 reflexed 
Upright stamens X reflexed stamens=? 
Whether these ratios are also representative of the laws that govern 
the transmission of types of flowers in Vitis rotundifolia is a question 
deemed worthy of investigation. 
The method of procedure naturally resolves itself into a study of 
the progeny of crosses between vines with different types of flowers. 
Since there are flowers of three distinct types, namely, staminate 
flowers with upright stamens, hermaphrodite flowers with upright 
stamens, and hermaphrodite flowers with reflexed stamens, we may 
expect to make five different combinations, as follows: 
1. Hermaphrodite flowers with reflexed stamens X staminate flowers 
(with upright stamens). 
2. Hermaphrodite flowers with reflexed stamens X hermaphrodite 
flowers with upright stamens. 
3. Hermaphrodite flowers with upright stamens X hermaphrodite 
flowers with upright stamens. 
4. Hermaphrodite flowers with reflexed stamens X hermaphrodite 
flowers with reflexed stamens. 
5. Hermaphrodite flowers with upright stamens X staminate flowers 
(with upright stamens). 
The types of flowers that were observed in the progeny resulting from 
the first cross (hermaphrodite flowers with reflexed stamens X stamin¬ 
ate flowers with upright stamens) has been partially discussed in Tech. 
Bui. FTo. 10 of this Station (5). Since that time more extensive and 
important data has been collected. 
The following table shows the results so far obtained: 
