166 
MUSCULAR WORK 
character and the amount of the work, and the metabolism measured at stated 
intervals after the work had ceased. The per minute total calorie output, 
the carbon-dioxide output, the oxygen consumption, and the respiratory 
quotient are included, together with the observations of the pulse- and respira- 
tion-rates. Naturally the measurements of the oxygen consumption have the 
greatest interest, since they form an admirable index of the total katabolism, 
and if corrections are made for the varying calorific equivalents of oxygen 
based upon the respiratory quotient as determined, the calorie production can 
be directly computed. One great difficulty with this type of experimenting 
is the fact that in order to determine accurately the respiratory quotient, 
exact determinations of the carbon dioxide are necessary, for if a resting 
period is begun immediately after the severe muscular work, abnormal 
results for carbon dioxide might be obtained as a result of retained carbon 
dioxide." 
Table 136. — Metabolism of subjects lying on the couch without food before and after work. 
[The subject of the experiment of Nov. 21, 1911, was H. L. H.; of Nov. 23, 1911, E. P. C; 
and of the remaining experiments, M. A. M. Figures in italics represent duration of 
work, heat equivalent of work, and the oxygen consumption per minute during the last 
period of work.] 
Date. 
1911. 
Nov. 21 
Nov. 23 
Dec. 15 
Dec. 20 
Dec. 21 
Dec. 22 
1912. 
Jan. 1 
Elapsed 
time 
from 
end of 
work to 
begin- 
ning of 
period. 
37 min. 
15 min. 
45 min. 
18 min. 
42 min. 
13 min. 
34 min. 
55 min. 
8 min. 
27 min. 
48 min. 
lh.9m. 
10 min. 
32 min. 
52 min. 
11 min. 
37 min. 
Carbon dioxide 
eliminated. 
Per 
min. 
200 
215 
200 
Increase 
after work. 
Amt. P. ct. 
-15 
21 
-10 
15 
Oxygen 
absorbed. 
Per 
min. 
Increase 
after work. 
Amt. P. ct. 
Heat 
output 
(computed) . 
Per 
min. 
c.c. c.c. I cals. 
240 .. I ... 1.18 
Work: 65 min. 73 cals. 
-7.0 I 259 I 19 I 7.9 I 1.23 
... 1211 I .. I ... I 1.01 
Work: 46 min. 4o cals. 
12.4 249 38 18.0 1.18 
-5.9 225 14 6.6 1.05 
In- 
crease 
after 
work. 
7.5 
4.6 
-1.0 
3.1 
250 1.20 
1 Vork: 75 min. 81 cals. 
255 5 2.0 1.24 
252 2 0.8 1.21 
226 
Work: 70 mm. 
1.10 
72 cals. 
258 
247 
252 
•■',2 
21 
20 
9.0 
2.0 
-1.5 
-3.5 
231 
Work: 73 min. 
14.2 
9.3 
11.5 
1.24 
1.18 
1.21 
250 
201 
217 
2-1(3 
28 
30 
16 
15 
16.0 
1.5 
2.1 
225 
Work: 78 min. 
12.1 
13.0 
6.9 
6.5 
1.12 
92 cals. 
1.26 
1.25 
1.18 
1.17 
1.09 
96 cals. 
25:i 
23S 
22S 
38 
u 
3 
12.4 
5.8 
1.3 
1.24 
1.15 
1.11 
233 ..I ... 1.14 
Work: 69 mm. 88 cals. 
7.7 268 35 15.0 1.30 
-1.9 246 13 5.6 1.19 
Res- 
pira- 
tory 
quo- 
tient, 
p. ct. 
0.89 
1,754 c.c. 
4.2 I 0.77 
I .80 
1,368 c.c. 
16.8 0.76 
4.0 .71 
80 
1,444 c.c. 
3.3 i 0.84 
.8 .80 
I .86 
1,344 c.c. 
12.7 
7.3 
10.0 
0.79 
.78 
.80 
1,256 c.c. 
12.5 
11.6 
5.4 
4.5 
0.84 
.78 
.79 
.78 
1,355 c.c. 
13.8 
5.5 
1.8 
0.89 
.83 
.87 
1,162 c.c. 
14.0 0.84 
4.4 .83 
Pulse-rate. 
Per 
min. 
Increase 
after work. 
Amt. P. ct 
12 
16.9 
16.0 
6.0 
25.0 
16.7 
11.7 
8.3 
16.7 
6.7 
3.3 
20.9 
1.5 
Res- 
pi ra- 
tion- 
rate 
per 
min- 
ute. 
a See discussion of this point, p. 148. 
