FLORA VITIENSIS. 255 
> “™y 
Other important differences have nob been observed, All this leaves no doubt that the two genera in 
question are indeed more closely related to each other than they are to any allied Order, and that they 
possess that degree of aflinity which justifies us in regarding them as belonging to one and the same 
Natural Order.” 
“ Under these circumstances, the establishment of the independent Natural Order Trochodendree, to 
be ranged with the other Polycarpee, appears to be perfectly justified, and as, in comparing them with the 
other Polycarpee, we have merely to keep in mind the Magnoliacee, Winteracee, and Schizandree, we ob- 
tain absolute differential characters, and a distinct habit. We may be allowed to lay considerable stress 
upon the want of floral envelopes, this character being constant also in Euptelia, We must also attach im- 
portance to the epitropous nature of the ovules (to which Agardh justly assigns great systematic value), 
meeting with it in Tpochodendree, and not finding it in the three Orders with which we havé compared them. 
If we add to these the differences which they respectively present, as, for instance, the want of stipules 
as contrasted with Magnoliacee (quite apart from the spathe-like development, the lower and higher leaf- 
formation of this Natural Order), and the want of oil-cells, so widely dittused in the Winteracee and Sehi- 
zandree, even in their foliage and bark,—if, finally, we take into consideration the distinct habit of the Z’re- 
chodendree as expressed by their serrated leaves, we shall have good evidence in fayour of the above con- 
clusions.” 
* It remains for me to express my thanks to Dr, Seemann, for kindly reminding me to examine the 
structure of the wood of Huptelia, with the view of ascertaining how far it agreed with that of Zrochoden- 
dron described in my paper. In complying with his wish, | found that Huptelia had the usual structure of 
deciduous woods,—a close dotted prosenchyma, traversed by numerous net-like vessels without any ob- 
servable peculiarities, and without any special characteristics agreeing with those of Trochodendron. We 
have here the same evident differences of anatomical structure as in the allied Winteracee between Llliefwm 
and Drimys, and an additional proof that botanical affinities and internal structure do not always go 
together.” 

I, Sponia, Comm. mss. ex Lamk. Dict. vol. iv. p. 188; Planch. in Ann, Se. Nat. dme Ser. 
t.x. p. 264. Flores polygami, 3-morphi. Fl. : Perianthium 5-partitum, lacints gestivatione sub- 
yalvato-induplicatis, marginibus tamen leviter quincunciatim-imbricatis, Antheras demum ex- 
sertze, loculis basi non conspicue gibbosis, rimis dehiscentiz introrsis, Yl. ¢; Perianthii fere mascult 
laciniis minus induplicato-valvatis et minus concavis; cetera J et >. Fl. 2: Perianthiilacim vix 
explanate, basi imbricate, apicibus non valvato-conniventibus, Bacca minuta, atylis 2 brevibus, 
plumoso-stigmatosis coronata, perianthio suffulta. Cotyledones falcate-conpaph cast path Geules, 
non corrugate,—Arbores inermes; foliis trinerviis ssepe canescenti- v, cinereo +pabescentiius, 
serratis ; eymis axillaribus in axillis foliorum adultornm et novellorum solitarits ¥ SeCsins, alits in 
ramulo codem masculis, aliis polygamis; foemineis seepius in ramulis propriis; pedicellis conspicue 
articulatis, articulis a sese invicem facile secedeutibus.—Dene. Deserip, Herb. Timor. p. 170. 
Besides the two species enumerated below, there 1s, 1n tropical Polynesia, S. discolor, Dene. (Celtis 
discolor Bronen., ©. orientalis, Sol. Prim. Fl. Ins. Pacif. (ined.) p. 345, et in Parkins, Drawnigs of Tatsit, 
Plants (ined )t 115 “ Ba-uhe,” Tahitensibus), from the Society Islands (Banks and Solander! Vorster ! 
.) t. 115, 
Barclay !). 
1. S. velutina, Planch. l.c, p. 3275 ramis novellis foliisque subtus molliter velutinis, in- 
: nte; foliis ovato-oblongis cuspidatis, basi leviter inzequali cordatis y. 
supra puncticulis ercberrimis asperis; cymis (3; 2, 3 que), breve 
petiolum sequantibus v. fere duplo superantibus, co-floris; or 
bus ¢ extus sparse pilosulis ; bacea ovata, apicem rarius pilis paucis albis Se ctr pi 
berrima,—Lakeba (Seemann ! n. 563). Also found on Uvea or Wallis Island (Gresfle! u, 57), 2 
in the East Indies, the Indian Archipelago, and China. | 
dumento in novellis splende 
rotundatis, margine serratis, 
pedunculatis v. subsessilibus, 
Planch. 1, ¢. p. 336; ramulis crassis petiolisque tenuiter sericeis, demum 
9 §. Andersonii, ee 8 
i stipulis dimidiato-oyato-lanceolatis, petiolum eequantibus; foliis ovato 
cinereo-pubescentibus 5 
