2021 SIMMONS ET AL.: NEW SPECIES OF MYOTIS 19 
DECIBELS (dB) 
90-70-50 -30 
150 
So 
oO 
FREQUENCY (kHz) 
3 

1050 1100 1150 1200 1250 1300 1350 1400 1450 
Time (ms) 
FIGURE 9. Spectrogram of echolocation calls emitted by the holotype Myotis nimbaensis upon initial release 
(FFT size 1024, Hanning window; sampling rate of 500 kHz). Color scale represents amplitude of sound in 
decibels (dB). 
In contrast to Myotis rufoniger and M. weberi, both M. bartelsi (FA = 53.4 mm; GLS = 20.42 
mm; ZB = 13.41 mm) and M. hermani (FA = 56.0-60.0 mm; GLS = 20.10-21.77 mm; ZB = 
13,4-14.10 mm) are somewhat larger than M. nimbaensis in most dimensions (Csorba et al., 
2014). As described by Csorba et al. (2014), M. bartelsi can be distinguished by having strong 
lambdoid crests (weakly developed in M. nimbaensis) and a P3 that is fully out of line with the 
rest of the toothrow and not visible in lateral view (partially out of line and visible in lateral 
view in M. nimbaensis). The forehead is also less clearly developed and strongly sloped in M. 
bartelsi compared to M. nimbaensis, and the rostrum appears somewhat shorter in M. bartelsi 
in lateral view. Myotis hermani as described by Csorba et al. (2014) can be distinguished by its 
overall very robust skull and exceptionally well-developed sagittal and lambdoid crests (more 
weakly developed in M. nimbaensis), very large upper canine with a basal area exceeding that 
of P4 (basal areas of C and P4 subequal in M. nimbaensis), minute P3 less than one quarter the 
size of P2 (P3 only slightly less than one half the size of P2 in M. nimbaensis), P3 fully out of 
line with the rest of the toothrow and not visible in lateral view (only partially out of line and 
visible in lateral view in M. nimbaensis), p3 with half the basal area of p2 (p3 is at least three 
quarters the basal area of p2 in M. nimbaensis), and p3 partly lingually displaced out of the 
toothrow (p3 fully in toothrow in M. nimbaensis). 
Asian Myotis (subgenus Chrysopteron) species exhibiting “formosus type” external mor- 
phology are M. formosus and M. rufopictus (Csorba et al., 2014), These species can be dis- 
tinguished from M. nimbaensis based on pelage color and banding. The dorsal fur of 
formosus-type species has individual hairs that have a very narrow brown base and are pale 
yellow distally for 80-100% of their length, or grade into a brown tip, with banding some- 
