2021 SIMMONS ET AL.: NEW SPECIES OF MYOTIS 15 

dorsal dorsal ventral iP ventral 
FIGURE 6. Uropatagium, foot, calcar, and flight membrane attachments in Myotis nimbaensis (AMNH 279589, 
holotype). A, Dorsal side of the uropatagium showing orangish brown skin with orange fur on the proximal 
1/3 of the surface of the membrane. B, Ventral side of uropatagium showing pale cream-colored fur on proxi- 
mal 1/5 of ventral membrane surface; this fur continues across the femur onto the proximal plagiopatagium 
in a narrow strip that does not extend past the knee. C, Dorsal view of distal leg, foot, calcar, and associated 
flight membranes. Note that the wing membrane (on right of the foot) attaches to the foot at the base of the 
first toe, and the calcar (on the left of the foot) is more than twice the length of the hind foot. D, Close-up of 
the dorsal surface of the foot showing sparse, long brown hairs on each toe. E, Close-up of the ventral side of 
foot and toes showing dark brown coloration. F, Ventral view of the hind leg and foot showing the relatively 
small foot size (foot length = 2/5 of tibia length) and the patterning of black membrane pigmentation near 
the leg and foot. 
oped sagittal crests (though again, our sample size is small). Upper premolars in M. tricolor are 
somewhat variable, but P3 is always less than half the basal area of P2 and is always shifted 
lingually as in M. nimbaensis. However, in many individuals the reduction in P3 is greater than 
in M. nimbaensis, and this tooth is completely excluded from the toothrow (only partly excluded 
in M. nimbaensis). The p3 of M. tricolor is one half to two thirds the basal area of p2, and p3 
is sometimes displaced lingually from the toothrow, compared to a slightly larger p3 that is at 
least three quarters the size of p2 and which is not displaced in M. nimbaensis. 
Another orange Myotis species from Africa is M. bocagii, which occurs across much of 
tropical Africa and is broadly sympatric with M. tricolor and M. welwitschii (Monadjem and 
Jacobs, 2017a; Patterson, 2019). Myotis bocagii can be easily distinguished from M. nimbaensis 
