2021 SIMMONS ET AL.: NEW SPECIES OF MYOTIS 
(1994) disagreed. Both of these authors overlooked a 1958 ruling by the International Com- 
mission on Zoological Nomenclature that fixed the gender of Myotis as masculine and placed 
the name as such on the Official List of Generic Names in Zoology (International Commission 
on Zoological Nomenclature, 1958). So in this case, nimbaensis is masculine. 
DiAGNosIs AND DEscrRIPTION: Myotis nimbaensis is diagnosed by a combination of the 
following characteristics: large size (FA 52.4—55,2 mm, mass 15.5-17.0 g; table 1) with females 
apparently somewhat larger than males; dorsal fur bright orange with strongly tricolored hairs 
(basal 1/3 of hair shaft black, middle 1/3 of shaft creamy white, distal 1/3 and tip bright orange 
to coppery red); ventral fur paler than dorsal fur, tricolored on belly (black base, buff-colored 
shaft that grades to orange at tip) grading to bicolored (lacking black base) on the sides near 
the wing membranes; ruff of brighter fur present around neck; pointed face with pale skin 
covered with orange fur, skin clearly visible through the fur around eye, mouth, and on ros- 
trum; no black spots present on face; pinna with a rounded tip, relatively long and reaching 
beyond tip of nose when laid forward; pinna with strong distal emargination; skin of pinna 
pale orange brown becoming slightly darker near the tip, no rim of black around ear; tragus 
lanceolate, slightly less than half the length of the pinna; thumb brown; wing membranes 
strongly dichromatic black and orange, with plagiopatagium and dactylopatagium mostly black 
with narrow bands of orange along the metacarpals, phalanges, and across the membrane 
behind the forearm and upper arm; black pigmentation extends medially nearly to body and 
posteromedially to tibia; membrane between metacarpals I and II pale orange; propatagium 
pale orange; uropatagium orangish brown with orange fur on the proximal 1/3 of the dorsal 
surface and pale cream-colored fur on proximal 1/5 of ventral surface; no black spots on pata- 
gia; no conspicuous gland present in plagiopatagium behind humerus; pale ventral fur extends 
onto proximal plagiopatagium in narrow strip along body, does not extend past knee; relatively 
small foot with length approximately 2/5 of tibia length; foot and toes brown, with sparse long 
brown hairs on the dorsal surface of each toe; wing membrane essentially naked except close 
to the body, where it has a sparse covering of cream colored hairs on the ventral surface; wing 
membrane attaches to foot at base of first toe; calcar long, more than twice the length of the 
hind foot, runs approximately 2/3 of the length of the uropatagium border; skull large for 
Myotis (GLS = 19,48-19.73 mm; CIL = 18.92-18.93 mm; CBL = 18.31-18.81 mm) with globu- 
lar braincase, well-defined sloping forehead, and elongated supraorbital region all contributing 
to the appearance of a clearly defined and elongate rostrum; rostrum with shallow medial 
depression in nasal region; sagittal crest moderately well developed in both sexes; lambdoid 
crest weakly developed in both sexes; dental formula 12/3, C1/1, P 3/3, M3/3 = 38; I1 and [2 
each with a well-developed posterior cusp; upper canine robust, crown height approximately 
2x that of P4, basal area (as seen in occlusal view) roughly equal to that of P4; anterior two 
premolars (P2 and P3) much smaller than last premolar (P4); P3 with less than half the basal 
area of P2 and shifted lingually to be partly excluded from the toothrow although still visible 
in lateral view; P4 large, sharply pointed, taller than M1; lower incisors each with 4 main cusps; 
i3 with distal accessory cuspules; p3 relatively large, approximately three quarters the basal area 
of p2, and fully in line in toothrow, not shifted lingually; lower molars myotodont. 
