

148 
rs in,its greater breadth and thickness in propor- 
tion to its length, and especially in the greater strength and outward extension of the 
condyle for the innermost of the anteriorly directed toes (Pl. CYT. figs. 5, 6, #) > the shaft 
of this composite bone is rather more twisted on its axis, with a Bret inclination of 
the stem of the condyle (#) backward, The * entocondylar cavity’ (ib. fig. 5, a) is somie- 
what deeper and larger than the ectocondylar one (4); the intercondylar tubercle (¢) is 
large, but little elevated. The ectocalcaneal process (ib, fig. 6, s) is a strong and pro- 
minent subquadrate plate of bone. The entocalcaneal process (ib, ib, 7) is, as usual in 
Raptores, of smaller size. The intervening calcaneal groove or channel (ib. ib. 2) is of 
great depth and width. Into the wide and deep antinterosseal depression (ib. fig. 5,2) open 
the entinterosseous and ectinterosseous canals, ‘The small hinder orifice of the latter 
remnant of the primitive interspace between the ecto- and mesometatarsal elements is 
shown in fig. 6, at m. The ectinterosscous groove is continued down a short way below 
this orifice. A strong tuberosity (fig. 5, 7) marks the insertion of the tendon of the 
‘tibialis anticus.’ The entogastrocnemial ridge (7) is continued from the entocalcaneal 
process nearly halfway down the shaft of the metatarsus. The ectogastrocnemial ridge 
(x) and the postinterosseous ridge (y) are well developed. The intermuscular ridges on 
the fore part of the shaft (fig. 5) are equally well marked; the entometatarsal ridge is 
shown at g, the ectometatarsal ridge at #. A long groove (0) for the ‘adductor digiti 
externi’ deepens as it leads to the foramen (p), through which the tendon of that small 
muscle glided to the interspace between the meso- and ectotrochleew. A strong osseous 
bridge (ib. fig. 6, q) divides the upper and hinder orifice (p') from the intertrochlear 
outlet of the tendinal canal, The depth and extent of the surface for the ligamentous 
attachment of the innermost and backwardly directed metatarsal (7) bespeak the strength 
of the toe opposing the forwardly directed digits (i & vz) in the grasping actions. The 
ectotrochlea (7v) is, transversely, rather narrower than usual relatively to the other 
trochlee; but it is of equal antero-posterior extent. The least circumference of the 
shaft of the metatarsal of Harpagornis moorei is 2 inches; the breadth of the two 
extremities and the length of the bone are shown in PI, CVI. 
Among the more characteristic evidences of the present extinct gigantic Raptorial 
are certain claw-bones (ungual phalanges). Assuming this ungual phalanx (Pl. CVIL. 
fig. 7) to correspond with the one which is commonly the largest in diurnal Raptores, 
viz. that which supports the back toe (digit 7), a second somewhat smaller claw-bone, 
discovered at the same time and place, and differing only in a slight inferiority of size, 
may well be a claw-bone of the toe ij. Subsequently a third ungual phalanx was 
discovered in another part of the Glenmark swamp, of rather less length than the 
second, but of equal size of basal articulation, and with it the penultimate phalanx of 
the same toe. On the assumption that the largest claw-bone (Pl. CVII. fig. 7) was that 
of the ‘hallux,’ or hind toe (é), it may be compared with the homologous bone in the 
Great Wedge-tailed Eagle of Australia (Aguila cuneicaudata of Brehm) or the Bold 
From that type the present fossil diffe 

