30 
Measurements (mm) and Weights (g) of Highland Specimens of Marmosops noctivagus, 
AMERICAN MUSEUM NOVITATES 
TABLE 10 
NO. 3466 
Including the Type Material of Nominal Taxa Originally Described as keaysi, dorothea, and yungasensis 









Peru: Puno Bolivia: La Paz 
AMNH AMNH AMNH AMNH BMNH AMNH AMNH AMNH 
160692 164726 16473» 16068¢ 1.6.7.794 72558¢ 2689365 268937! 
Sex female female male male female female male male 
HBL 132 127 143 151 130 136 130 124 
LT 185 178 175 204 158 169 168 163 
HF 17 19 22 22 17 17 19 19 
Ear — — — — 25 — 24 24 
CBL 35.7 — 35.1 39.5 34.2 35.6 34.1 34.5 
NB 4.4 3.9 4.8 4.6 43 4.7 4.1 4.2 
LIB 6.8 6.5 7.2 75 5.9 6.6 6.4 6.3 
ZB 17.7 17.3 18.6 20.0 17.5 18.3 17.5 17.6 
PL 20.7 20.0 20.5 22.7 19.2 20.5 19.6 20.0 
PB 10.7 10.8 10.9 11.6 10.9 10.6 10.4 10.6 
MTR 14.5 14.6 14.2 16.0 13.9 13.6 13.8 14.4 
LM 6.9 7.2 6.7 h2 6.7 6.3 6.7 6.9 
MI-3 5.9 6.1 5.8 6.2 5.8 5.6 5.8 5.8 
WM4 2.3 2.5 2.4 2.6 2.4 2.2 2.5 2.5 
Weight — — — _— — — 37 37 
aParatype of keaysi. 
bTopotype of keaysi. 
‘Holotype of keaysi. 
dHolotype of dorothea. 
eHolotype of yungasensis. 
fFrom Chijchijpa; AMNH 268936 was sequenced by Mustrangi and Patton (1997). 
synonym of dorothea (= M. noctivagus in 
our usage; see above), M. ocellatus is an un- 
ambiguously diagnosable taxon that more 
closely resembles M. impavidus in morphol- 
ogy and mtDNA sequences (tables 4, 6). 
Externally, Marmosops ocellatus is dis- 
tinctively paler than other Bolivian species, 
with sandy grayish-brown dorsal fur and 
self-cream underparts that lack distinct lat- 
eral zones of gray-based hairs. The metapo- 
dials are covered dorsally with pale (whitish) 
hairs that do not contrast abruptly in color 
with the digits. The tail is distinctly bicolored 
(dark above, pale below) and _ particolored 
(paler distally than proximally), such that the 
distal one-third or more of the organ is com- 
pletely pale in most specimens. Most exam- 
ined males have no trace of a gular gland, 
the only exception being the holotype (an un- 
usually large specimen; table 11), in which 
the gland appears to be present. The lateral 
carpal tubercles of adult males are bulbous 
or knoblike, and the scrotum is pale (with 
whitish fur and unpigmented skin). Three 
parous adult female specimens prepared in 
the field by L.LH. Emmons (LHE 1468, 1596, 
1808) each had 6-1-6 = 13 mammae, of 
which the anteriormost two pairs appear to 
be pectoral in digital images that we exam- 
ined. The supraorbital margins of the skull 
are rounded, lacking distinct beads even in 
the largest specimens, and a shallow postor- 
bital constriction is dorsally visible. Palatine 
fenestrae are consistently present, the upper 
canine lacks accessory cusps, and the audi- 
tory bullae are moderately large. 
Marmosops ocellatus differs from M. im- 
pavidus by its paler and grayer dorsal col- 
oration, absence of distinct lateral zones of 
gray-based ventral hairs, possession of pec- 
toral mammae, and more distinctly bi- and 
particolored tail. In addition, the palate is 
more extensively fenestrated and the audito- 
ry bullae are much larger in M. ocellatus 
than in M. impavidus. Although the differ- 
ence between exemplar sequences of these 
