2004 
VOSS ET AL.: BOLIVIAN MARMOSOPS 3 
TABLE 1 
Valid Species and Synonyms Currently Referred to the Genus Marmosops* 



Taxa Types? 
M. bishopi (Pine, 1981)¢ USNM 393535 
M. cracens (Handley and Gordon, 1979) USNM 418503 
M. dorothea (Thomas, 1911)4 BMNH 1.6.7.79 
Synonyms: ocellatus (Tate, 1931)¢ BMNH 26.1.5.25 
yungasensis (Tate, 1931)4 AMNH 72558 
M. fuscatus (Thomas, 1896) BMNH 96.11.1.6 
Synonyms: carri (J. A. Allen and Chapman, 1897) AMNH 7314/5922 
perfuscus (Thomas, 1924) BMNH 23.11.13.18 
M. handleyi (Pine, 1981) FMNH 69838 
M., impavidus (Tschudi, 1845) 
Synonyms: albiventris (Tate, 1931)4 
Not known to exist 
USNM 194378 
caucae (Thomas, 1900) BMNH 99.9.6.51 
celicae (Anthony, 1922) AMNH 47182 
madescens (Osgood, 1913) FMNH 19689 
oroensis (Anthony, 1922) AMNH 47180 
sobrinus (Thomas, 1913) BMNH 13.10.24.70 
ucayalensis (Tate, 1931) AMNH 78952 
M. incanus (Lund, 1840) 
Synonyms: bahiensis (Tate, 1931) 
[ZMUC 223, 224; L.14, L.15}f 
BMNH 3.9.5.137 
scapulatus (Burmeister, 1856) ZMB 2330 
M. invictus (Goldman, 1912) USNM 178708 
M. juninensis (Tate, 1931)¢ AMNH 63864 
M. neblina Gardner, 19902 USNM 560735 
M. noctivagus (Tschudi, 1845) ZMB 33754 
Synonyms: collega (Thomas, 1920) BMNH 20.7.14.40 
keaysi (J. A. Allen, 1900) AMNH 16068 
leucastrus (Thomas, 1927b) 
lugendus (Thomas, 1927a) 
BMNH 26.4.1.102 
BMNH 27.1.1.178 
neglectus (Osgood, 1915) FMNH 19636 
politus (Cabrera, 1913) MNCN 797 
purui (Miller, 1913) USNM 105543 
stollei (Miranda-Ribeiro, 1936) MNRJ 1267: 
M. paulensis (Tate, 1931) FMNH 26576 
M. parvidens (Tate, 1931) FMNH 18545 
M. pinheiroi (Pine, 1981)¢ USNM 461459 
Synonym: woodalli (Pine, 1981) USNM 393532 



aSynonyms (including “subspecies”) are listed under each species recognized as valid by Gardner (1993) except as noted 
otherwise below. Full bibliographic information for all original descriptions is provided in the References section. 
bHolotypes by original designation unless noted otherwise. 
cListed as a synonym of M. parvidens by Gardner (1993) but recognized as a valid species by Voss et al. (2001). 
dSynonymized with M. noctivagus in this report. 
eConsidered to be a valid species in this report. 
fSyntypes. Tate (1933: 164-165) claimed that only two specimens in Copenhagen (ZMUC 223 and 224) were actually collect- 
ed by Lund and are therefore eligible to be “co-types” (= syntypes), but two uncataloged skulls (labelled L.14 and L.15) are also 
part of Lund’s original material and therefore have the same nomenclatural status. Because ZMUC 224 is the only specimen that 
consists of both a skin and cranial material (223 is a skin only), it would be a logical choice for lectotype. 
8Listed as a synonym of M. impavidus by Gardner (1993) but recognized as a valid species by Patton et al. (2000). 
hA “co-type” (= syntype) according to Tate (1933: 153). 
iSee Langguth et al. (1997). 
jListed as a synonym of M. incanus by Gardner (1993) but recognized as a valid species by Mustrangi and Patton (1997). 
