160 FIELD COLUMBIAN MuUSEUM—GEOLOGY, VOL. II. 
ward at this place, but that it curved forward more after the way of birds, 
a position compatible with the upright attitude. The whole structure 
of the neck, the elongated vertebre, the transverse centra, and lateral 
exapophysial articulations, the absence of transverse processes, all remind 
one of the neck in the cryptodire testudinates, where the neck is capable 
of much and effective antero-posterior flexion. 
The articulation of the head of the femur, with the basal plane of its 
convexity nearly at right angles to the long axis of the bone, and the 
posterior position of the acetabulum, carry conviction to me that the 
femora could not have been brought parallel in the same direction with- 
out dislocation from. the sockets, whatever attitude the animal may 
have assumed. ‘The articulation of the distal end of the tibia further- 
more shows that the metatarsus could not have been extended to a right 
angle; that is, P/eranodon could not have been plantigrade. The greater 
extension must have been between the metatarsus and phalanges; in other 
words, the creature must have been digitigrade in ambulation. The flat- 
tened proximal ends of the metatarsals indicate a compact and closely 
united foot, and the bones have been so found in specimens of Pteranodon. 
The toes in /feranodon, and doubtless in this genus also, were practically 
clawless, and the outer toes were much the longer. The animal was 
incapable of seizing or holding with the feet. 
Nor could a very great flexion at the knee have been possible, as I 
think the figures of the femur of P¢eranodon ingens will indicate. If 
the animal was quadrupedal, it must have been in a crawling attitude, 
with both legs and arms widely extended. 
APATOMERUS MIRUS, GEN. ET. SP. NOV. 
In the University of Kansas Quarterly, vol. iii, p. 3, I described and 
figured a remarkable bone from the Lower Cretaceous of Clark County, 
Kansas, which I hesitatingly referred to some crocodile-like animal, - 
because I was at a total loss where else to place it, my knowledge of the 
pterodactyl anatomy then being less than at present. The figure and 
description are reproduced in vol. iv of the University Geological Sur- 
vey of Kansas, p. go, as follows: ‘‘The upper end of a femur found in 
the same region appears to belong to the same kind of an animal, as 
does the vertebra described above. The shape is not unlike that of a 
human femur, with the trochanters evidently small and placed much 
below the level of the head. The neck is stout, the head gently convex, 
with an angular border. The shaft below the trochanters is somewhat 
flattened from before backward, but becomes more cylindrical below. The 
shaft is hollow, with firm walls not more than one-third of an inch in 
thickness, The portion preserved theasures 210 mm.” 
