VULGARIS L. AND IN PH. MULTIFLORUS WILLD 105 
causes the colourdevelopment in the eye itself. Thus eight different 
genotypic combinations may be realized: (1) PTE, (2) P Te, (3) 
PtE, (4) Pte, (5) pTE, (6) pTe, (7) ptE, and (8) pte. Num- 
bers 1 and 2 are totally coloured individuals (E is supposed to 
be hypostatic to T); eyed beans occur in number 3, and the other 
combinations 4—8 have white, colourless seeds. Besides the possi- 
bility remains, that different factors (E’, E"), are to be accepted 
for the distinctive types of pigmentextension. For there are races 
of beans, that show within narrow limits of fluctuations, the pigment 
restrained to a very small spot around the navel, while in other 
races the pigmentextension occupies a third or more of the whole 
seedcoat. And this pigmentextension is an essential character of 
the race. ; 
By the way it may be mentioned, that in Vigna sinensis also 
a factor for totality has been discovered (SPILLMAN 1913, MANN 
1914, HARLAND 1919), leaving in its absence only a part of the 
seedcoat coloured, while the rest is white. : 
The further factors, as discovered in researches that have been 
published till now, are those concerning the various colours and 
colourintensities, so much characteristic for the seedcoats of 
Phaseolus vulgaris. TSCHERMAK in his earlier papers already has 
made an effort to divide both groups of coloured individuals: the 
marbled. contained three colourdifferences (black, violet and brown) 
and these three groups occurred in the proportions of 9:3:4 (92 
blackmarbled : 33 violetmarbled : 38 brownmarbled, thus in more 
precised proportion 9: 3.23 : 3.72); the selfcoloured ones however 
showed the reverse proportion of 4 blacks:3 violets:9 brown 
(39 black:27 violet: 101 brown, or 4: 2.77: 10.36). These results 
of TSCHERMAK’S paper may be tabulated in this manner: 
pigmented 3 : white 1 
marbled 1 : selfcoloured 1 
Bl. M. 9:V.M. 3: Br. M. 4. BĲ, Se 42¥.8.-32Bric8: Oor 
A more detailed grouping of the colours met with a great many 
difficulties and uncertainties concerning the conclusions for the 
genotypic differences : a few proportions only (brownmarbled: yellow- 
marbled and full-black : brownish-black) showed to follow a simple 
mendelian relation 3:1. (30:8 and 29:10); in other cases the 
proportions of segregation were less clear. For explaining these 
