132 SIRKS, THE COLOURFACTORS OF THE SEEDCOAT IN PHASEOLUS 
If truly the pollenparent in this case has been a white-seeded 
individual, this white-seeded plant will probably have been an 
A-individual, that missed all other colourfactors or in which all 
other colourfactors were in a kryptomeric state. The formula of 
the hybrid will perhaps have been AABb GsLIZz; a further 
research of the F;-families will give here the correct explanation. 
12c. Cross oF BRUINE X WITTE. 
The black-yellow-brown marbled pheenotype is of course not 
the only one, that can take its origin after the cross Bruine X Witte. 
Other spontaneous crossings were found, though in remarkable 
less numbers. So for example reddish brown-yellowbrown marbled 
individuals; their segregation was of course less complicated than 
that of the blackmarbled F;-plants. Five phaenotypical groups only 
were observed in these segregations: I redbrown marbled, II 
yellow-brown-marbled, III redbrown selfcoloured, IV yellow-brown 
selfcoloured and V white. The frequencies of these groups in the 
Fa-generations that were grown and in some F3-families, that were 
the posterities of redbrown marbled Fo-individuals, coming from 
a cross Bruine X Witte, and segregating along the same lines, are 
tabulated in the following table: 








Type TE HS | IVe M | Total 
1044 Spont. hybr. from Bruine 8 Zeit 4 7 32 
1047 „ 5 A rs 4 1 6 1 5 17 
1048 F; after spont. hybr. from Bruine | 20 | 7 | 19 6 | 19 71 
1049 , » b 5 À je LS dels SAT 49 
1050252 0 5 > 4 & 13.1, 001 14 4 | 12 59 
61 | 21 | 68 | 20 | 54 | 224 
Other F3-families, derived from redbrown marbled Fo-plants 
segregated in still less phaenotypical groups; these may also be 
given here in a table: 5 | : 
Cf table page 133 
The whole segregation indicates a trihybrid nature of the F,-plants, 
the formula of which can be written as AaBbGGLI: its descendants 
segregate thus in five phaenotypical groups: I ABbGL, II ABbGI, 
IT ABB (or bb) GL, IV ABB (or bb) Gl and V white a-plants. 
ED EN ay WR 
