AND SPECIES HYBRIDS WITHIN THE GENUS SACCHARUM 141 
This makes it highly probable, that the sluggards are the univalent 
chromosomes of the nuclear plate and that the chromosomes near the 
poles have been derived from the gemini. As we shall see in the last 
chapter, other investigators have found analogous anaphases in the 
case of plants showing in the prophase of the division of the PMC the 
presence of gemini besides unpaired chromosomes. They also conclu- 
ded that the sluggards were the univalent chromosomes. Of these chro- 
mosomes they could observe the longitudinal splitting. That really 
longitudinal splitting occurs in the case of sugarcane is not certain, on 
account of the smallness of the chromosomes ; it is however probable. 

Fig. 38. Fig. 39. 
Fig. 38. Anaphase of the heterotype division of Green German New Guinea (irre- 
- gular division) x 2300. 
Fig. 39. Anaphase of the heterotype division of G. G. N. G. (irregular division) 
x 2300. 
Fig. 38 shows an anaphase of G. G. N. G. in which all the univalent 
chromosomes are still near the aequator, some of them split already. 
In Fig. 39 we see the chromosome-groupes lying somewhat nearer to 
the poles, while nearly all the sluggards are split, but their halves — in 
general clearly smaller than the chromosomes near the aequator —, 
are still in the proximity of the aequator. 
In fig. 40 one sees that the chromosomes derived from the gemini 
have almost reached the poles while the unpaired ones are distributed 
all along the spindle. In part the latter are split into halves; others 
show no splitting as yet, while it seeems that some fall to pieces. 
