

A NEW DOMINANT COLOR PATTERN AND COMBINATIONS 
PAS bhEED TRUEVIN THE GROUSE LOCUSTS 4) 
by 
ROBERT K. NABOURS, Manhattan, Kansas, U.S.A. 
Among the grouse locusts (Tettigidae) there are many conspicuous 
dominant color patterns. Something is known about the factor rela- 
tions of forty-one of these elementary patterns in three species of as 
many genera. Thirty-two have been reported, Paratettix texanus (Na- 
bours ’14, °17), Apotettix eurycephalus (Nabours ’19), and Tettigidea 
parvipennis (Bellamy ’17). Only one of these elementary, patterns is 
definitely known to have originated in the laboratory. 
This new pattern, in P. texanus, was first reported as possibly a case 
of linkage of some kind (Nabours ’17(d), p. 48, Mating 472B IS), but con- 
siderable subsequent experience with those that are unmistakably lin- 
ked has changed that view. It has not yet been discovered in nature. It 
is Strikingly distinct and certainly germinal, having maintained its in- 
dividuality eight years, through twenty-seven generations, and nume- 
rous combinations (Plate, Sm/Sm. Originally designated IS IS). It se- 
gregates alternatively from the original pattern (Plate, S/S) from which 
it mutated, as well as from eleven others of the multiple allelomorphs 
with which it has been combined (Nabours ’17, B, C, D, E, F, H, I, J, 
N, P and Q). The individuals bearing the factor for this new pattern 
are at least as vigorous as any of those found in the wild state. Speci- 
mens of it were placed in the field in 1921, and again in 1922, and al- 
though not yet recovered, it is believed that they will survive as well as 
any. 
Whatever the cause, a profound change has occurred in the factor for 

1) Paper No. 67. Department of Zoology, Kansas State Agricultural College 
Experiment Station. 
