516 THE CYTOLOGY OF THE SUGARCANE 
with 68 chromosomes in the haploid phase. All the same, 2364 POJ 
possesses the same number of chromosomes as the Cheribon-Kassoer 
hybrids. 
It is therefore not impossible, that 2364 POJ has arisen from the fer- 
tilisation of an eggcell of roo POJ which contained 40 chromosomes. 
The eggcell which, after fertilisation, produced G g2 must however 
have possessed more than 40 chromosomes. Probably therefore — as 
in the case of other hybrids between species differing in chromosome- 
number — chromosomes are eliminated in unequal numbers in the re- 
duction-division of the megaspore-mothercells of 100 POJ, so that egg- 
cells with different chromosome-numbers are formed. 
In the case of EK 28 we must therefore also count with the possibili- 
ty that this cane may have originated from the fertilisation of an egg- 
cell of zoo POJ with 40 chromosomes by a sperm-nucleus of EK2 with 
an equal number of chromosomes; but the cytology gives no certain 
answer to the question whether EK r8 has been derived from zoo POJ 
or not. a? 
At the utmost we might consider the great irregularity in the reduc- 
tion-division as a slight indication of such an origin, because the des- 
cendants of Loethers (one of which is roo POJ) show manyirregulari- 
ties in the reduction-division, but other kinds of cane show this also. 
THE FERTILITY OF THE INFLORESCENCES OF SUGARCANE 
In the first part attention was fixed already on the frequency of ir- 
regularities in the reduction-division of the sugarcane and all kinds of 
sugarcane described in this part showed these irregularities. In the 
first part we also pointed out the possibility of a connection between 
the course of the reduction- division and fertility. As an example we 
mentioned, that Ardjoeno-cane which is rather fertile in the male line 
usually shows a complete reduction-division, while in the case of 
Bandjermasin mtam, on the contrary, which possesses incomplete re- 
duction-division only, the pollenformation is very poor, while that 
‚which is formed, is moreover but slightly fertile. To the existence or 
non-existence of a connection between the way in which the reduction- 
division occurs and the degree of fertility one can however conclude 
only after having made a large number of observations running over 
