338 THE POPULAR SCIENCE MONTHLY 
zoic, but have since been in a state of degeneracy, their present repre- 
sentatives being few in number and of a depauperate character. The 
application of this law throughout the enormously lengthy periods re- 
quired for the evolution of existing species, has led to the survival of 
some of the most ancient types until the present day; to the absolute 
obliteration of others which at one time gained great prominence; and 
to the gradual dying out of yet others, some of which are now found in 
the last stages of their existence. But through the entire course of 
change, the evolution of higher and yet higher forms has been the most 
conspicuous fact. Furthermore, it is undoubtedly true that the general 
course of evolution is in progress to-day as in the past, since all the 
potentialities of such evolution exist now as always, though conditioned 
by the fact that owing to continued changes in the physical character 
of the earth’s atmosphere as well as of its crust, the possibilities of evolu- 
tion are steadily diminishing and will eventually cease. 
There is one direction in which paleobotany gives well-defined as- 
surance that the evidence derived from existing species leads to correct 
conclusions. In tracing the succession of types, we are led to the belief 
that there is no direct sequence. Conterminous evolution is in accord 
with neither theory nor ascertained facts, and it is, therefore, impossible 
to conceive of a figure which shall in any way represent a single and 
unbroken line of succession. If paleontology teaches us anything, it is 
that each great phylum, as well as its various subdivisions, finally reaches 
its culmination in a terminal member from which no further evolution 
is possible. But that from some inferior member, possessing high 
potentialities, a side line of development arises. There is thus, in the . 
early life of each member of the series, a certain recapitulation of 
ancestral characters. This conception of a continuance of the main 
line of descent through a succession of lateral members is both logical 
and fully in accord with the evidence derived from both recent and 
extinct forms of plant life, as well as with our present theory of 
evolution. 
PALEONTOLOGY AND ISOLATION 
By Dr. JOHN M. CLARKE 
STATH MUSEUM, ALBANY, N. Y. 
HE notion of isolation as a factor in variation, as I am using the 
term, is that of geographic separation exclusively, the concep- 
tion expressed most clearly by Wallace, Moritz Wagner and Jordan. 
I take it that while this influence has been carefully estimated in the 
geographical distribution of living species, it has not often been ex- 
pressed in its own terms in the analysis of extinct faunas. With in- 
creasing accuracy in the record of ancient continental lines and bar- 
