122 
Fishery Bulletin 119(2-3) 
Acknowledgments 
The study was supported by the state assignment to the 
Shirshov Institute of Oceanology of the Russian Academy 
of Sciences (no. 0149-2019-0009) and financially supported 
by the Russian Science Foundation (no. 19-14-00026). 
In memoriam 
Sergei A. Evseenko was a world-renowned scientist, doctor 
of biological sciences, and head of the Laboratory of Oceanic 
Ichthyofauna of the Shirshov Institute of Oceanology, 
Russian Academy of Sciences. The main scientific interests 
of S. Evseenko included the taxonomy and ecology of the 
early stages of development of fish. He developed an eco- 
logical classification of flounder species, described for the 
first time the larvae of several dozen species of oceanic fish, 
and revealed the patterns of distribution of ichthyoplank- 
ton in the upwelling areas of the eastern Pacific Ocean 
and frontal zones of the World Ocean. He described several 
species and genera of marine fish. S. Evseenko described a 
new family of armless flounders of the Southern Ocean— 
the family Achiropsettidae—and clarified the morphology, 
anatomy, individual development, and evolution of this new 
family. In collaboration with N. Parin and E. Vasilyeva, he 
published a collective monograph, Fish of the Russian Seas: 
Annotated Catalogue. Overall, S. Evseenko published about 
170 scientific works. On 15 October 2020, he died suddenly. 
S. Evseenko was working on this manuscript just before his 
death. Everyone who is fortunate enough to have known 
this wonderful person and outstanding scientist will keep 
a bright memory of him. 
Literature cited 
Becker, V. E. 
1983. Myctophid fishes of the World Ocean, 248 p. Moscow, 
Russia. [In Russian.] 
Becker, V. E., and S. A. Evseenko. 
1986. Distribution of mesopelagic fishes and biogeographic 
boundaries in the southern part of the Pacific Ocean in 
January—February 1985. J. Ichthyol. 26:890—901. [In Russian.] 
Bolshakova, Ya. Yu., and S. A. Evseenko. 
2015. Larvae of the lanternfish Lampanyctus intricarius 
(Myctophidae) from the southwestern Pacific Ocean. J. 
Ichthyol. 55:596-600. 
2016. On the species composition of ichthyoplankton of the 
Walvis Ridge (South Atlantic). J. Ichthyol. 56:848-860. 
2020. Ichthyoplankton of the central North Atlantic Ocean: lar- 
val development of lanternfish species (Pisces: Myctophidae). 
Fish. Bull. 118:135-144. 
Denton, J.S.S. 
2014. Seven-locus molecular phylogeny of Myctophiformes 
(Teleostei; Scopelomorpha) highlights the utility of the 
order for studies of deep-sea evolution. Mol. Phylogenet. 
Evol. 76:270-292. 
Fricke, R., W. N. Eschmeyer, and R. Van Der Laan (eds.). 
2020. Eschmeyer’s catalog of fishes: genera, species, refer- 
ences. [Available from website, accessed November 2020.] 
Hulley, P. A. 
1981. Results of the research cruises of FRV ‘Walther Herwig’ 
to South America: 58. Family Myctophidae (Osteichthyes, 
Myctophiformes). Archiv. Fischereiwiss. 31(1):1-303. 
Kobyliansky, S. G. 
2016. Anatomy of fishes of genus Lampanyctus (sensu lato), its 
taxonomic structure, and status in the Lampanyctini (Mycto- 
phidae) tribe system. J. Ichthyol. 56:181—199. 
Martin, R. P., E. E. Olson, M. G. Girard, W. L. Smith, and M. P. Davis. 
2018. Light in the darkness: new perspective on lanternfish 
relationships and classification using genomic and morpho- 
logical data. Mol. Phylogenet. Evol. 121:71-85. 
McGinnis, R. F. 
1982. Biogeography of lanternfishes (Myctophidae) south 
of 30°S. Antarct. Res. Ser. 35, 110 p. Am. Geophys. Union, 
Washington D.C. 
Miller, J. M., W. Watson, and J. M. Leis. 
1979. An atlas of nearshore marine fish larvae of the Hawaiian 
Islands. Sea Grant Misc. Rep. UNIHI-Seagrant-MR-80-02, 
179 p. Univ. Hawaii Sea Grant Coll. Prog., Honolulu, HI. 
Moser, H. G., and E. H. Ahlstrom. 
1974. Role of larval stages in systematic investigations of 
marine teleosts: the Myctophidae, a case study. Fish. Bull. 
72:391-413. 
1996. Myctophidae: lanternfishes. In The early stages of 
fishes in the California Current region (H. G. Moser, ed.), 
p. 387-475. CalCOFI Atlas 33. 
Moser, H. G., and W. Watson. 
2001. Preliminary guide to the identification of the early life 
history stages of Myctophiform fishes of the western central 
Atlantic. NOAA Tech. Memo. NMFS-SEFSC-453, 118 p. 
Moser, H. G., E. H Ahlstrom, and J. R. Paxton. 
1984. Myctophidae: development. In Ontogeny and systemat- 
ics of fishes. Based on an international symposium dedicated 
to the memory of Elbert Halvor Ahlstrom; La Jolla, 15-18 
August 1983 (H. G. Moser, W. J. Richards, D. M. Cohen, 
M. P. Fahay, A. W. Kendall Jr., and S. L. Richardson, eds.), 
p. 218-239. Am. Soc. Ichthyol. Herpetol., Spec. Publ. 1. 
Olivar, M. P. 
1985. Ictioplancton del Atlantico sudoriental. Ph.D. diss., 710 p. 
Univ. Barcelona, Barcelona, Spain. [Available from website.] 
Olivar, M. P., and L. E. Beckley. 
1997. Larval development of Lampanyctus species (Pisces: Myc- 
tophidae) from the southwestern Indian Ocean, and species 
groups based on larval characters. Bull. Mar. Sci. 60:47-65. 
Pertseva-Ostroumova, T. A. 
1918. Mediterranean Scopelidae (Saurus, Aulopus, Cholor- 
phthalmus, and Myctophum). In Report of the Danish ocean- 
ographic expeditions 1908-1910 to the Mediterranean and 
adjacent seas, vol. 2 (Biol.) (A.7), 154 p. Hgst, Copenhagen, 
Denmark. 
1964. Some morphological characteristics of myctophid lar- 
vae (Myctophidae, Pisces). Tr. Inst. Okeanol. 73:76—92. [In 
Russian.] 
Taylor, W. R., and G. C. Van Dyke. 
1985. Revised procedures for staining and clearing small 
fishes and other vertebrates for bone and cartilage study. 
Cybium 9:107-119. 
Wisner, R. L. 
1976. The taxonomy and distribution of lanternfishes (family 
Myctophidae) of the eastern Pacific Ocean. NORDA Rep. 3, 
229 p. Navy Ocean Res. Dev. Act., Bay St. Louis, MS. 
Zahuranec, B. J. 
2000. Zoogeography and systematics of the lanternfishes of 
the genus Nannobrachium (Myctophidae: Lampanyctini). 
Smithson. Contrib. Zool. 607, 69 p. [Available from website.] 
