U.S. Department 
of Commerce 
Gina M. Raimondo 
Secretary 
National Oceanic 
and Atmospheric 
Administration 
Richard W. Spinrad 
Administrator 
National Marine 
Fisheries Service 
Janet Coit 
Assistant Administrator 
for Fisheries 
Fishery Bulletin (ISSN 0090-0656) 
is published quarterly by the Scientific 
Publications Office, National Marine 
Fisheries Service, NOAA, 7600 Sand 
Point Way NE, Seattle, WA 98115-0070. 
Although the contents of this publica- 
tion have not been copyrighted and may 
be reprinted entirely, reference to source 
is appreciated. 
The Secretary of Commerce has deter- 
mined that the publication of this peri- 
odical is necessary according to law for 
the transaction of public business of this 
Department. Use of funds for printing of 
this periodical has been approved by the 
Director of the Office of Management and 
Budget. 
For Sale by the Superintendent of 
Documents, U.S. Government Publishing 
Office, Washington, DC 20401. Subscrip- 
tion price per year: $32.00 domestic and 
$44.80 foreign. Cost per single issue: 
$19.00 domestic and $26.60 foreign. See 
back for order form. 
risnery 
Bulletin 
Scientific Editor 
José |. Castro 
National Marine Fisheries Service, NOAA 
263 13th Avenue South 
Saint Petersburg, Florida 33701 
Managing Editor 
Kathryn Dennis 
Scientific Publications Office 
National Marine Fisheries Service, NOAA =\= 
1845 Wasp Boulevard, Building 176 
Honolulu, Hawaii 96818 
Associate Editor 
Cara Mayo 
Scientific Publications Office 
National Marine Fisheries Service, NOAA 
7600 Sand Point Way NE 
Seattle, Washington 98115-0070 
Editorial Board 
Henry L. Bart Jr. 
Katherine E. Bemis 
Matthew D. Campbell 
William B. Driggers III 
Tulane University, New Orleans, Louisiana 
National Systematics Laboratory, Washington, DC 
National Marine Fisheries Service, Pascagoula, Mississippi 
National Marine Fisheries Service, Pascagoula, Mississippi 
Gretchen L. Grammer South Australian Research and Development Institute, Adelaide, Australia 
Richard Langton National Marine Fisheries Service, Walpole, Maine (retired) 
J. Femando Marquez-Farias Universidad Autonoma de Sinaloa, Mazatlan, México 
Richard S. McBride National Marine Fisheries Service, Woods Hole, Massachusetts 
Richard L. Merrick National Marine Fisheries Service, Falmouth, Massachusetts (retired) 
Richard D. Methot National Marine Fisheries Service, Seattle, Washington 
Lisa J. Natanson National Marine Fisheries Service, Narragansett, Rhode Island (retired) 
Mark S. Peterson University of Southern Mississippi, Ocean Springs, Mississippi 
André E. Punt University of Washington, Seattle, Washington 
Joseph M. Quattro University of South Carolina, Columbia, South Carolina 
John F. Walter III National Marine Fisheries Service, Miami, Florida 
Fishery Bulletin website: https://spo.nmfs.noaa.gov/fb.htm 
Fishery Bulletin carries original research reports on investigations in fishery science, 
engineering, and economics. It began as the Bulletin of the United States Fish Commission 
in 1881; it became the Bulletin of the Bureau of Fisheries in 1904 and the Fishery Bulletin 
of the U.S. Fish and Wildlife Service in 1941. Separates were issued as documents through vol- 
ume 46; the last document was no. 1103. Beginning with volume 47 in 1931 and continuing 
through volume 62 in 1963, each separate appeared as a numbered bulletin. A new system 
began in 1963, with volume 63, in which papers are bound together in a single issue. Beginning 
with volume 70, number 1, January 1972, Fishery Bulletin became a periodical, issued quarterly. In 
this form, it is available by subscription from the Superintendent of Documents, U.S. Government 
Publishing Office, Washington, DC 20401. It is also available free in limited numbers to libraries, 
research institutions, and state and federal agencies and in exchange for other scientific publications. 
