SUBFAMILY ERIOSOMATINAE 347 
Subfamily Eriosomatinae Kirkaldy 
Kirkaldy, 1905:418. 
Characters. Vertex convex; frontal tubercles lacking. Eye in fundatrix 
three-facetted, more or less rudimentary in summer aptera, compound in alata. 
Wax glands present. Secondary sensoria on antenna transverse or annular. 
Unguis shorter than base of VI (except in fundatrigenia of Mordvilkoja). Hairs 
inconspicuous (except in Asiphum). Cornicle rudimentary or absent. Cauda 
semilunar, inconspicuous. Fore wing with media once-forked or simple. Rostrum 
four-segmented, i.e. with V indistinct and rudimentary. Sexuales degenerate, 
small and without rostrum; ovipara laying but a single egg and hind tibia with- 
out sensorialike structures; male apterous and without secondary sensoria on 
antenna. 
Living concealed in galls, pseudo galls, underground on roots, in ants nests 
or exposed on plants but covered with cottony secretion. 
Key to Tribes of Eriosomatinae 
1. Wax glands composed of a row of facets surrounding a central space; cornicle 
distinct, a pore usually on slight mammiform base. (p. 347) -.........:sc-tecccseeseeseeee Eriosomatini 
Wax glands not composed of row of facets around central space; cornicle 
MTICLASLINGCE WIN CLE V7 atl tN OTOP ASEM Ewe eectesepeee cree ete ccsesa ce reccreetnscasaceccencece sh cacsczdset enc assdsonieeceassenactevsusdvess 2 
2. Secondary sensoria suboval to narrow and transverse. Living in galls or 
pseudo galls on primary host (except Prociphilus venafuscus Patch, on bark), 
Onroots or stems-on- secondary hostia (pe 354) kcsc-cecs--cececce-cccccepeecesntéado-sbossseceeaccaagsacsess Pemphigini 
Secondary sensoria oval to suboval. Living subterranean and with ants; 
Primal vans ce UNO Wits 1M NU) Se Asel (Pe OZ2) ees cccsacessedcreseeasecutcsdeeedececcelqecssonseiecesovelSiacicusacasbe Fordini 
Tribe Eriosomatini Baker 
Baker, 1920a:65. 
Characters. Secondary sensoria on III of alata usually annular. Otherwise 
as given in key and under subfamily. 
Key to Genera of Eriosomatini 
1. Farsi distinctly two-segmented in all forms; hind wing with 2 cross-veins.........-.......-.----- 2 
... Tarsi with suture between I and II indistinct or lacking in aptera; hind 
WINPUWIEhNIONe CrOSS-V.C1NcOniISiIM Dp 1 Os (Pir OA 7.) sacsecs ses tosis sess waaerem cena c5eecue-taceas gen aanuscacarenepincnacete Colopha 
2. Fundatrix with antenna five-segmented; cornicle not apparent (p. 349)................ Eriosoma 
Fundatrix with antenna four-segmented; cornicle apparent (p. 353)... Gobiashia 
Genus Colopha Monell 
Monell, 1877:102; Baker, 1920a:66; Maxson, 1923:316. 
Characters. Antenna four-segmented in fundatrix, five-segmented in sum- 
mer vivipara, six-segmented in alate vivipara; secondary sensoria in alata 
annular. Tarsal I not distinctly separated in aptera. Media of fore wing simple 
or once-forked; cubitus absent or rudimentary in hind wing. Cornicle on 
mammiform base or lacking. Living subterranean or in galls. 
Genotype (monotypical), Byrsocrypta ulmicola Fitch. 
Colopha appears synonymous with Tetraneura Hartig if Aphis ulmi Geof- 
froy (n.n. T. ulmifoliae Baker) (the species that Hartig described but named 
“Aphis ulmi L. ?”) is considered genotype. However, this procedure is contrary 
to Opinion 65, in accordance with which Aphis ulmi L. would be genotype which 
would place the genus Tetraneura with Eriosoma. 
Genotype Colopha ulmicola (Fitch) 
The Cockscomb Aphid of Elm 
Byrsocrypta ulmicola Fitch, 1859:843. 
Tetraneura graminis Monell, 1882:16; Patch, 1910b:196 and 208; Gillette and Palmer, 
1943 :214. 
Colopha graminis, Hottes and Frison, 1931a:348. 
Colopha ulmicola, Baker, 1920a:66; Cutright, 1925:200 (synonymy); Hottes and Frison, 
1931a:349. 
