PARTULA CLARA. 65 
their relationship is indicated by attenuation of the shell as a whole, by the thinness 
of the lip, by the plain offspring of one of their number, and by the fact that most of 
them have no counterparts among the banded individuals found further south. 
Hence they can not be regarded as wanderers from a locality where a distinct stock 
of striped snails originated. 
Partula clara minor var. nov.—Orofere to Tiamao Valleys. 
After leaving Aoua Valley, one encounters a variety that maintains the same 
characters throughout all the valleys to Tiamao, when again there is an abrupt 
change. The area of its occupation includes Orofere, Atitara, Vaitupa, Vaipuaril, 
and Tiamao Valleys, of which the last is in the southern sector of the island. The 
shells of this region are much like the examples taken from the area to the east, 
which was originally inhabited by clara, as described by Garrett; they differ from 
the latter in some details, however, notably in their extreme breadth and in the 
brevity of the aperture. It is justifiable for our purposes to separate them as a 
distinct variety to be designated minor (figs. 37 to 42, plate 20, and fig. 1, plate 
Diatalew23)). 
The classes of “lighter” (figs. 38 and 41, plate 20) and “darker” (fig. 37, plate 20) 
shells are not as clearly distinguishable as heretofore, although extreme individuals 
of both kinds are found in almost all of the valleys now under consideration. All 
of the Atitara specimens are practically intermediate, while in Vaitupa there is a 
well-marked group of “medium” individuals (fig. 39, plate 20), dark types being 
absent. In Tiamao, all of the unbanded specimens are extremely light in ground- 
color. The banded shells (figs. 40 and 42, plate 20; fig. 1, plate 21) are characteristic. 
Here fuller data are at hand regarding the heredity of the color characters, 
although records for Orofere and Atitara could not be obtained. The facts prove 
that the color types do not breed true in all cases; 6 “light” Vaitupa adults 
contained 15 well-grown embryonic young, all of which were “light”; 5 “medium” 
adults contained 5 “light” and 4 “medium” young, although it must be added that 
the young of any one parent were alike in color value. In Vaipuarii, 5 adult 
records are available; 1 light adult bore 4 similar young, while 4 dark adults 
contained 5 dark young. All of the unbanded Tiamao individuals were light, and 
they bore 17 equally light young. In brief, a type tends to produce its like, but 
exceptions occur, as in the case of the Vaitupa series. 
Passing to the banded specimens, and the fixity of their color peculiarity, it 
must be restated that all of the young of unbanded parents were similarly devoid of 
the rotating marks. The banded individual from Vaitupa gave no data, but the 
2 specimens from Tiamao were both gravid; the first of these bore 2 banded 
young, and the second had 1 similar embryo. Scant as the data are, they are 
at least strongly indicative of a fixed heritable nature of the banded character. 
Turning now to the statistical characteristics of the variety under consideration 
(table 23), we find, as stated earlier, that the shells are about as long as those of var. 
parva, but they are much wider and consequently much stouter. The aperture is 
also broad in proportion to its length, but otherwise it is not distinctive. 
