72 VARIATION, DISTRIBUTION, AND EVOLUTION OF THE GENUS PARTULA. 
contracted apertures (fig. 24, plate 21). There is a progressive change in virtually 
all of the shell characters in passing to Tenaire and further to Titaviri (table 26). 
The shells become shorter and wider and relatively stouter, while the apertures 
become somewhat longer, much wider, and relatively broader; the same sequence 
appears in the proportion of aperture length to shell length. 
The collections from the valleys of Taiarapu present no features of special 
interest, aside from their diversity. Of the valleys in the eastern quadrant, only 
Paraura afforded a fair collection; this series is in general more like the southern 
shells than like those of the peninsula. The resemblance is emphasized when the 
comparison is made by regions (table 26), when all of the smaller valley collections 
are taken into account as well as those of the representative valleys. 
As there are no banded or dark-colored shells belonging to this variety, the 
statistics of heredity are of little interest beyond the fact that neither of the two 
general color-classes lacking among the adults is represented among the embryonic 
but advanced young; of the last, there are Iog in all. 
Partula clara var. prima (?).—Papenoo Valley. 
The last valley on the east and north which is inhabited by clara is Papenoo, 
a very large element in the series of valleys situated in the northern quadrant. The 
discovery of this species in a locality so remote from its original headquarters is less 
remarkable than the fact that the shells found in Papenoo are entirely different from 
marmorata, whose range, we have seen, extends the whole distance from this point to 
the former boundaries of the species. From the bushes of the level valley floor 
5 living specimens were secured in 1906 about 2 miles inward from the coast; of 
these, 2 were light and unbanded (fig. 30, plate 21), while 3 were banded like the 
Vaihiria examples (fig. 31, plate 21). In 1908 and again in 1909 this valley was 
visited in order to secure additional specimens of clara, if possible, as well as further 
information regarding the separate species of stolida Pease, reported by Garrett as 
occurring only in this valley. In spite of prolonged search, however, no additional 
clara were obtained. Apparently the 1906 examples belonged to a small localized 
colony existing in the lowlands. 
The resemblance of this colony to the Vaihiria shells is marked especially by 
the character of the banded shells; the unstriped light examples are also like prima 
and differ absolutely from marmorata. In statistical characters, furthermore (table 
27), there appears to be a closer relation to var. prima than to any other regional 
variety. 
TaBLeE 27.—Shell characters of five Partula clara from Papenoo Valley. 
SS ay 
Character. Mean value. Standard deviation. 
Shell length imm yen ens Grae tetas nicks seen 15.7600 .1513 0.5018 .1071 
wid thyme Be ee aso ee ees «eR 9.1400 .0800 . 2653+ .0566 
PDUROVOLOM BLO, Oe CANE. cdooudoooosouns00000 57.9000+ .4514 1.4966 .3194 
Aperturemlensth mma rrelrereecrteteisierisieieiae 8.9000 .0539 .1788= .0381 
Widths mM yc vierccichetetemitasienisttetae ys 6.5400+ .0451 1496+ .0319 
Proportions spemcents. eerie 73.5000 .1907 6324+ .1349 
Length aperture + length shell, proportions, p. ct..) 56.1000 .1477 -4898 = .1045 
