PARTULA OTAHEITANA. 1@ fe) 
from which the others are absent. To avoid establishing too many taxonomic 
grades, we will speak of this whole part of P. otaheitana as the “sinistral series of the 
south and west,” and will give the status of primary varieties to its three constituent 
elements, under their original descriptive names. While each variety is much 
diversified within its subordinate area, and, indeed, within the confines of a single 
valley, the shells of the whole series agree incertainfundamental characters. ‘Theyare 
typically sinistral, although dextral sports occur here and there, at times attaining 
considerable numbers; they are various shades of brown in color, displaying the rich 
yellow and red of types like rwbescens only in certain definitely localized associations; 
revolving brown bands are developed in a considerable number of individuals in 
nearly every colony, while certain unbanded color-types recur in nearly all valleys. 
Partula otaheitana sinistrorsa occurs in certain southwestern valleys of Taiarapu 
and in the southern sections of Tahiti nui as far west as Taharua Valley; sinistralis 
occupies the territory from Teohu to Vaipuarii Valleys, inclusive; finally crassa 
inhabits the western sector from Vaitupa to Taapuna Valleys. 
H. The circuit of the island is completed by P. otaheitana lignaria Pease, which 
inhabits only Tipaerui Valley, adjacent to Fautaua. ‘This variety is almost exclu- 
sively dextral, relatively small, brown and never red in color, and it is usually 
strigated; its casual resemblance to P. o. affinis disappears on closer study. Plain 
and banded shells constitute the two color-classes. In some parts of its valley, 
reversed shells of both color-divisions occur infrequently. On the whole, this 
variety articulates with P. 0. otaheitana toward the east, and with hitherto unde- 
scribed relatives which exist in the neighboring valleys of the western sector. 
PARTULA OTAHEITANA OTAHEITANA VAR. NOV. 
GENERAL STATEMENT. 
The members of this species that live in Fautaua Valley are first to be taken 
up, both because the original specimens described by Bruguiére seem to have 
belonged to this colony and also because they form as a group the most generalized 
division of the whole otaheitana series, similar to the original stock from which all 
of the other geographical and taxonomic sections have originated. On account of 
its primacy in both senses, this colony will be designated as P. otaheitana otaheitana. 
Bruguiére’s original description, quoted in Pilsbry, is as follows: 
“This species is fluviatile and was discovered during the same voyage as the preceding 
[Capt. Cook’s] in the brooks of the island Otahiti. The shell is sinistral, oblong, oval, very 
thick, not more than 10 to 11 lines in length by about 6 in diameter. It is composed of 
5 whorls, united exteriorly, and as much swollen as those of the austral Bulimus, but coiled 
in a direction contrary to that of most shells, that is to say, the right side of its animal is 
toward the left [sic]. The spire is conic and terminates in a pointed summit. The sutures 
resemble those of the preceding species [P. faba]. The aperture is semi-oval, oblique, 
rounded at the base, only a third longer than wide, and shorter by a line than half the shell. 
Outer lip arcuate, much reflexed and recurved to the exterior, flat and projecting outside. 
The inner (parietal) lip is very thin and furnished with a tooth in the middle, found only 
in old shells, the young having an inconspicuous callus in its place. The columella is 
