272 VARIATION, DISTRIBUTION, AND EVOLUTION OF THE GENUS PARTULA. 
In fecundity, as in structure, the small series of banded snails is different from 
the others to a remarkable degree (table 227), for it shows a relatively high percent- 
age of gravid adults and a high fertility among them. Although it is conceivable 
that one group might resume breeding activity before the other, the eggs are about 4 
to 1 of the young in both series; so that no difference in the t7me of breeding can be 
invoked to account for the discrepancies. 
The embryonic young are all sinistral and devoid of the band. Apparently 
in this valley, as before, cestata shells do not display the stripe until late in life. 
TaBLE 227.—Partula otaheitana crassa. Atitara Valley. Fecundity. 
No. of | Percent} No. of | No. of Total Average Average 
gravid. | gravid. eggs. young. | contents. | for gravid. for all. 
Series. 
Plain, sin 
Banded, sin 
All, sin 
OROFERE VALLEY. 
In Garrett’s descriptive account, the type locality of crassa is given as Faahuaite 
Valley, which from intrinsic evidence is to be identified with the area now called 
Orofere on the charts, and Faarahi colloquially. The last name means simply 
“large valley”? and is used because the adjoining ravines on both sides are much 
smaller. The identification is based upon the existence of typical crassa in Orofere, 
and also upon the fact that this valley is the only large element lying between the 
range of sinistralis and Punaauia Valley, the home of the original Partula nodosa; 
Garrett mentions only the greater clefts, and speaks of Faahuaite as followed by 
Punaruu (Punaauia). As the name ‘‘Faahuaite” was not known to the local chiefs 
consulted by me, we must conclude either that Garrett’s information was incorrect 
or that his memory was at fault. In any case, there is no doubt whatever that 
the valley herein called Orofere is the place specified by the earlier author as the 
habitat of crassa in his time. 
Collections were secured in 1907 and 1908. On the first occasion, snails were 
taken in the lower valley up to a point about 3 miles from the coast and a mile 
beyond the point where Partule were first met at a barometric level of 200 feet. 
In 1908 a deeper journey was made, but as in the first year Partule were found in 
scattered situations and in small numbers. The two series were alike only in the 
relative preponderance of phza shells (95.5 per cent and 95.2 per cent, respectively) 
and in the lack of their sharp differentiation into striata and confluens; typical repre- 
sentatives of these are found, but the intergrades are so numerous as to render 
impossible a classification like that of Vaitupa and Atitara shells. The illustrations 
(plate 33, figs. 22 to 28) show several representatives of this class, typical and 
aberrant. Cestata shells are few, 6 being taken in 1907 and 3 in 1908; usually the 
distinctive markings are faint, but in the specimen of plate 33, figure 30, two bands 
are sharply defined. The shell of plate 33, figure 29, is bicolored upon the last 
whorl through the spread of the median band over the entire lower, or right, area. 
