52 WF. Ponder, D.J. Colgan, Т. Terzis, S.A. Clark, А.С. Miller 
Mannosephosphate isomerase MPI 5.3.1.8 1 1 
Peptidases: Phe-Pro substrate PPR 34.11 1 1 
: Leu-Ala substrate LAL 3.4.11 1 
: Leu-Leu-Gly substrate LGG 34.11 2 
Phosphoglucomutase PGM 5.4.2.2 2 2 
6-Phosphogluconate dehydrog. 6-PGDH 1.1.1.44 1 1 
Sorbitol dehydrogenase SDH 1.1.1.14 2 2 
Triosephosphate isomerase ТРІ 5.3.1.1 1 1 
UDP glucose pyrophosphorylase UDPG 2.1.1.9 1 1 
Results 
Taxonomy 
Three new species in two genera, one new, are described below, as are the electrophoretic results 
confirming the status of these species. 
Dalhousia n. gen. 
Type species: Dalhousia globosa n. sp. 
Diagnosis: Hydrobiids with simple, smooth, imperforate, broadly ovate to conic shell, with simple 
aperture and lacking external varix. Operculum with weakly to moderately-developed pegs on inner 
side and radula with two laterally placed basal cusps on central teeth. Female genital system with 
single globular, posteriorly-located medium-sized sperm sac and posteriorly elongated coiled oviduct. 
Male with long, tapering (whip-like) penis. Stomach with horn-like process on upper edge of 
posterior chamber and lacking obvious caecum. 
Description: Shell (Figs 2А-С, F-H, 3-5) simple, smooth, imperforate, broadly ovate to conic, with 
simple oval aperture and lacking external varix. Protoconch (Fig. 6A,B) of about 1.5 whorls; 
sculptured with irregular, shallow pits. Operculum (Fig. 7A-E) thin, flat, with white smear and 
weakly to moderately-developed pegs on inner side. Body heavily pigmented (mainly black), cephalic 
tentacles with narrow, median-dorsal unpigmented line (corresponding to line of dorsal cilia) and 
narrow unpigmented zone at base of tentacles. Mid-dorsal strip of cilia rather weakly-developed and 
set in a narrow, shallow groove (Fig. 8B,C); three more strongly developed cilial strips ventrally, two 
ventro-lateral and one mid-ventral (Fig. 8A,B). Snout (Fig. 9A-D) with little or no cilia dorsally: 
some cilial tufts latero-ventrally. Labial area weakly papillate, papillae with very short cilia-like 
processes (Fig. 9D). 
Anatomy generally similar to Fonscochlea (Ponder et al., 1989). Columellar muscle short and 
wide. Ctenidium large, with numerous broadly triangular filaments, posterior end usually slightly 
overlapping anterior edge of pericardium. Osphradium oval, short, behind middle of ctenidium. 
Hypobranchial gland very narrow, weakly to moderately developed. Rectum straight or slightly 
arched in pallial cavity, with longitudinally orientated faecal pellets and, usually, partially 
overlapping pallial oviduct glands. Radula (Fig. 10) with 2 pairs of basal cusps on central teeth; 
otherwise typical of family. Stomach (Fig. 11) with characteristic "horn"-like process on upper edge 
of posterior chamber with caecum rudimentary or absent. Renal organ protruding partly into pallial 
roof, renal opening usually with white lips, renal gland longer than wide. Prostate gland (Fig 
12A,B) broadly oval, much higher (i.e. dorso-ventral dimension) than thick, mostly visceral, with 
pallial vas deferens opening near posterior pallial wall. Penis (Figs 12С, 13) long, tapering, whip" 
like, with pointed distal end. Penial duct (Fig. 12C) undulating in small, moderately expanded basal 
section, straight in long distal part. Female reproductive system similar to Fonscochlea but with only 
a single globular sperm sac (Fig. 14, sps). Coiled oviduct (Fig. 14A,B, co) with simple, U-shaped 
proximal part, long posterior extension, wrapped around sperm sac in S-shaped coil; proximal part 
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