62 W.F. Ponder, D.J. Colgan, Т. Terzis, S.A. Clark, А.С. Miller 
Dimensions of specimens from paratype series 
Females, М-11 
SL SW AL AW BW СУ ТУУ 
Minimum 2.68 1.86 1.35 1.29 2.19 0.12 2.6 
Maximum 3.70 2.60 1.88 1.64 3.01 0.19 3.6 
Mean 3.16 2.28 1.64 1.49 2.60 0.16 3.1 
S.D. 0.27 0.19 0.13 0.10 0.21 0.02 0.27 
Males, N=9 
SL SW AL AW BW CV TW 
Minimum 2.57 1.89 1.32 1.25 2.14 0.14 2.8 
Maximum 3.45 2.53 1.79 1.59 2.77 0.19 3.6 
Mean 3.00 2.16 1.55 1.41 2.45 0.16 BI 
S.D. 0.30 0.22 0.17 0.11 0.22 0.02 0.20 
Shell dimensions of 50 topotypes are given in Appendix 2, along with those of additional populations 
of this species. 
Operculum (Fig. 7С-Е). Ovate, pale yellow, thin, with eccentric nucleus, with 0—5 small, weak 
pegs on white smear on inner side. 
Statistics of 20 opercula from the type series. 
Females, М-11 
OL OW OS PL PH 
Minimum 1.11 0.70 0.30 0.12 0.05 
Maximum 1.53 1.12 0.53 0.32 0.13 
Mean 1.32 0.92 0.39 0.22 0.09 
S.D. 0.12 0.11 0.07 0.06 0.03 
Males, №9 
OL OW OS PL PH 
Minimum 1.11 0.77 0.26 0.17 0.07 
Maximum 1.38 1.05 0.55 0.27 0.14 
Mean 1.24 0.88 0.36 0.22 0.10 
S.D. 0.09 0.10 0.09 0.03 0.03 
Radula (Fig. 10A,E). As for D. globosa but central teeth usually with 3 (range 3-4) cusps on either 
side of the median cusp. 
Head-foot. Cephalic tentacles pigmented dorsally, with or without narrow unpigmented mid-dorsal 
stripe; snout and dorsal neck also pigmented. Ciliation (Fig. 8A,B, 9B-D, 13C) as for generic 
description. Foot and opercular lobes pigmented; pallial roof and visceral coil darkly pigmented, 
often partly mottled. 
Anatomy. Osphradium unpigmented, located at posterior quarter of ctenidium, narrowly oval, 0.22- 
0.40 mm in length. Ctenidium with 23-25 broadly triangular filaments, apices central to towards 
right. maximum width 0.32-0.48 mm, posterior-most filament overlapping pericaridum (rarely 
abutting or 2 filaments overlapping). Rectum straight in females, weakly arched in males. Renal 
organ extending 1/3-1/2 length of renal gland into pallial roof; renal gland 0.51-0.72 mm in length, 
0.26-0.34 mm in width. Pericardium extending 1/3 of length into pallial roof. Stomach (Fig. 11A) 
with style sac 0.46-0.62 mm in length, stomach proper 0.49-0.62. mm in length, caecum absent 10 
very small (up to 0.05 mm long). 
Testis of 1.8-2.0 whorls; seminal vesicle visible externally and lies between digestive gland and 
testis for only 0.1-0.3 whorls behind stomach, convolute over stomach. Prostate gland (Fig. 12B) 
