896 STYLIDIACEAE. [Donatia. 
A. DownatinaE. Corolla polypetalous. Stamens 2-3, free .. 1. DONATTIA. 
B. STYLIDIBAR. Corolla gamopetalous. Stamens 2, connate with the style into a 
gynostemium or “column.” 
Leaves densely imbricated, forming hard bright-green patches. 
Flowers sessile among the leaves, Capsule turbinate .. 2. PHYLLACHNE. 
Stems slender, branched. Scapes longer than the leaves. 
Capsule ovoid or oblong = § ® .. 3} FORSTERA. 
Stems short. Leaves tufted, squarrose, subulate. Scapes 
shorter than the leaves ‘ - ts .. 4, OREOSTYLIDIUM. 
1. DONATIA Forst. 17 7£ 
Small densely tufted herbs, forming hard compact masses. Leaves | 
densely imbricated, linear, coriaceous, quite entire. Flowers terminal, 
solitary. Calyx-tube adnate to the ovary, obconic; lobes 3-7, equal or 
unequal. Petals 5-10, linear-oblong to ovate. Stamens 2-3, inserted on 
the middle of an epigynous disc, and adnate to the base of the styles ; fila- 
ments subulate or filiform; anthers didymous, extrorse. Ovary inferior, 
2-3-celled ; styles 2-3, short and thick or subulate, recurved ; stigmas simple 
or capitellate ; ovules numerous, affixed to placentas that are pendulous 
from the inner angles of the cells. Capsule turbinate, indehiscent, 2—3-celled. 
Seeds few in each cell, pendulous, obliquely ovoid; testa membranous ; 
albumen fleshy ; embryo small, remote from the hilum. 
A genus of 2 species, 1 found in New Zealand and Tasmania, the other 
(D. fascicularis Forst.) a native of Fuegia. Its exact systematic position has been 
much discussed. It was referred to the Saxifragaceae by Hooker, who, however, also 
pointed out its affinity with the Styl:diaceae, with which it agrees in the stamens being 
placed on the centre of an epigynous disc, in the extrorse anthers, and in the placenta- 
tion. It was removed to that family by Baron Mueller (‘‘ Knovo Giornale Botanico 
Italiano,”’ xi, July, 1879). Mildbraed in the ‘‘ Pflanzenreich ’’ (Heft 35) also retains 
it in the Stylidiaceae ; but Skottsberg (“The Plant World,” May, 1915) suggests that it 
Don aT IN CERES ROREY Plat would . 16 (14180137 
1. D. novae-zelandiae Hook. f. Fl. Nov. Zel. 1 (1853) 81, t. 20. — Stems 
short, 1-3in. high, densely tufted, forming broad compact masses in 
mountain-bogs. Leaves very numerous, imbricited in many series and 
clothing the entire stem and branches, erect, appressed, 4 in. long, subacute, 
shining, velnless, very thick and coriaceous, villous at the base. Flowers 
din. diam., sunk amongst the uppermost leaves. Calyx-lobes. 5, ovate, 
acute. Petals 5, quite free, ovate-oblong, obtuse, thick and fleshy. Stamens 
2, styles 2, short and thick, recurved. Capsule 4 in. long.—Handb. N.Z. Fl. 
(1864) 58; Benth. Fl. Austral. 1 (1864) 450; F£. Muell. Fragm. vii (1873) 
41; Cheesem. Man. N.Z. Fl. (1906) 184; Muildbraed in Pflanzenr. Heft 35 
(1908) 20, fig. 7, D-F. (tAoue. So+ifP? — 1+ 4). 
Norta Istanp: Mount Holdsworth, Tararua Range, W. Townson! SovutH 
Istanp: Not uncommon in alpine bogs throughout. Srewarr Isranp: Petrie / 
T. Kirk ! Cockayne. Mostly between 3000 and 5000 ft., but descends almost to sea- 
level on Stewart Island. December—March. 
& 9. PHYLLACHNE Forst. '~ “¢ 
Densely tufted perfectly glabrous moss-like plants, forming hard and 
compact flat or convex masses in alpine localities. Leaves small, closely 
imbricating. Flowers sessile among the leaves at the tips of the branches, 
monoecious or polygamo-dioecious. Calyx-tube obconic ; lobes 5-9, equal 
