69 
A NEW GENUS OF CRASSULACEAE 
little pointed at the apex, connate at the base, slightly crinkled, micro- 
scopically papillose, bent in the upper portion towards the centre of 
the flower, about 1 mm. long, persistent, in bud deep red, in the open 
flower paler, later marcescent and dirty white. Stamens alternating 
with the petals, attached to their basal connate portion, half their 
length, bent sharply inwards through the chinks between the posterior 
portions of the carpels. Filaments from a broadish base subulate. 
Anthers brown, dorsifixed, oblong. Pollen yellow. The ovary fills the 
interior of the flower touching the petals. It is 4-chambered, 4-sulcate, 
depressed, almost flat on the upper surface and here microscopically 
papillose. In the centre, where the four grooves meet, there is a deep 
depression, close to which there are the four extremely minute free 
styles each of which bears an extremely small oblong stigma. The ovary 
is pale green except round the bases of the styles, where there is a 
little suffusion of red. The “ squamae” are a little longer than the height 
of the ovary, white, narrowly linear, slightly broadened towards the 
apex. It seemed to me that they are absent in some flowers, but on this 
point I could not quite make sure. The ripe fruits have not been seen 
yet. Fruits which were evidently nearly ripe were very slightly larger 
than the ovary in the open flower. They had a very thin pericarp which 
could easily be detached all round near the base of the ovary. It is, 
therefore, most likely that the fruit is a circumscissile capsule. The seeds 
are fairly numerous, oblong. They are attached to comparatively long 
funicles. 
In conclusion a few words on the placentation which is somewhat 
peculiar. Unfortunately we know too little of the placentation of other 
crassulaceous plants, especially of the small Crassulae which I have 
united under the section Tillaeoideae, to express an opinion whether 
this placentation is unique in the order. In each chamber of the ovary 
there are two quite separate placentas on the upper portion of the 
lateral walls near the inner angles of each chamber. Their shape and 
position will be seen from the accompanying figures (13, 14). Each 
bears numerous ovules. (PI. hi, a.) 
-2 
