REPORT OF THE DIRECTOR, 1922 63 
The fragmentary preservation of the pinnules would seem to indi- 
cate that they were of a rather delicate nature. There is no evidence 
of a midrib. The pinnules are set comparatively far apart so that 
the general effect of the foliage in the case of Eospermatop- 
teris is much less dense (figure 4) than is the case with Ly gin- 
opteris, Heterangium and others of the seed ferns. 
Seeds. The seeds of Eospermatopteris bear a strong 
external resemblance to those of Lyginopteris oldhamia, 
called Lagenostoma lomaxi™” before the relation between 
the two was known, and to other Lyginopterid seeds. The character 
of the seeds is well shown on plate 10, figures 5-12, plate 11, figures 
I-5, and in text figure 6. On some of the slabs the seeds are very 
abundant but on none of the specimens are any sterile laminae found 
in conection with the seeds, though branches bearing pinnules occur 
frequently in the same layers. The seeds were probably borne near 
the tip of the frond or the tips of some of the pinnae, with complete 
abortion of the sterile laminae; or perhaps there was modification 
of the bilobed pinnules. At any rate, the seeds are borne in pairs 
at the ends of forked branchlets. The ultimate division bearing the 
pair of seeds may be as short as a quarter of an inch, or less, or may 
be almost an inch long. In a number of cases when the seeds are 
borne close to the dichotomy, the arrangement of the fruits is rather 
crowded, and there is an appearance given of more than one seed 
being borne on a stalk. Sometimes the dichotomies are such a short 
distance apart as to bring, frequently two, sometimes three, pairs of 
seeds close together, giving a clustered effect to the seeds. The 
seeds, as in the case of Lagenostoma, are inclosed in an 
outer husk or cupule which completely envelops the seed. In some 
cases the cupule appears to come to a point at the top (plate 1o, 
figures 9, 10); in a few cases, some of which are figured here on 
plate 10, figures 7, 8, the cupules appear to be lobed as in Lagen- 
ostoma lomaxiandL.sinclairi (figure 5). This may, since 
it is infrequent, be only an accident of preservation; or it may be 
that the majority of these fruiting bodies are in an immature condi- 
tion. Some specimens in a very immature condition have been 
found. A few specimens show the impression of what appears to 
be a vascular bundle running up through the main branchlet, forking 
at the dichotomy and continuing up each arm of the dichotomy 
” Oliver, F. W. and Scott, D. H., Proc. Roy. Soc., 1903, 71:477-81; Phil. 
Trans. Roy. Soc. (B), 1905, 197:193-247, pls. 4-10; Scott, D. H., Studies 
in ae Botany, pt II, 1900, p. 386-96; Seward, A. C., Fossil Plants, 1917, 
3°55—04. 
