154 WISE 
In his book on New Zealand Coleoptera, G.V. Hudson (1934) gave a description 
of the Cicindela perhispida elytral markings and previous locality records. The 
systematic index included C. perhispida Broun and C. campbelli Broun. 
W. Horn (1936) again recorded the three known northern coastal forms of 
Cicindela but placed campbelli as a subspecies of perhispida. He gave additional 
locality records, repeated the Hokianga-Marsden Point confusion, and mentioned the 
Hokianga specimen of C. brevilunata (Horn 1926b) now in his collection from the 
Broun collection in the British Museum. 
In a note under the heading of Cicindela brevilunata, A.E, Brookes (1944) 
pointed out that this species is an east coast species and that C. perhispida is a west 
coast species. He queried the earlier confusion of localities, particularly a statement by 
Horn (1936), and made other comments. 
E. Rivalier (1963, 1964) described a new genus Neocicindela for all New Zealand 
species. Subsequently, C.M.C. Brouerius van Nidek (1965) listed the species of 
Neocicindela Rivalier, 1963 and included N. brevilunata and N.perhispida., For the 
latter, he recognised the subspecies perhispida perhispida and perhispida campbelli, 
and described a third subspecies perhispida giveni from Spirits Bay (which is on the far 
northern coast of Northland). 
In a catalogue of types held in the former Deutsches Entomologisches Institut, H. 
Débler (1973) listed “brevilunata Horn, 1926 (Cicindela)” with the three syntypes 
recorded by Horn (1926b) and the two of these in the DEI collection. Also listed was 
“brouni Horn, 1893 (Cicindela)” (a variety of perhispida synonymised with campbelli 
by Horn 1896) with one syntype in the DEI collection. The Deutsches Entomologisches 
Institut of Berlin was moved to Eberswalde in 1964 and was re-named as the Abteilung 
Taxonomie der Insekten of the Institut fiir Pflanzenschultzforschung, Eberswalde, 
D.D.R. (East Germany), in 1970. 
RECENT COLLECTIONS, SAND-DUNES AND SAND 
Coastal sand-dune collections 
Localities collected by the author since the 1960s (see Fig. 11) are listed in order 
from south to north up the west coast and southwards down the east coast in Table I. 
Specimen elytral marks (see Figs. 12-51) and colours are compared with visible sand 
colours and composition. Colours given are comparative, there is a range of 
intervening shades, There is also a wide range of variation in the elytral dark marks 
and colour pattern, particularly through the west coast to far north series (the 
campbelli-perhispida-giveni patterns). 
No secondary sexual characters have been noticed. Elytral colour patterns of 
males and females appear to fall within the overall range of variation. Larger 
individuals (often females) may give the impression of thicker dark marks but this is 
a matter of size, The key comparison is between the dark marks and the pale colour 
markings on each specimen. The thicker and thinner dark marks recorded here are 
visible on individuals of both sexes and all sizes. 
