1915.] Matthew and Granger, Lower Eocene Wasatch and Wind River Faunas. 93 
Pachyena ossifraga OSBORN (& WorTMAN) 1892, Bull. Am. Mus. Nat. Hist., 
Vol. IV, p. 112, fig. 11B; Matrrarw 1909, Mem. A. M. N. H., vol. IX, p. 489, 491, 
text fig. 91. 
Pachyena intermedia WORTMAN 1899, tbid., Vol. XII, p. 147; Matrrurw 1909, /. c. 
(type only). 
Type, U.S. Nat. Mus. No. 1096, an upper molar (m!, 1.) from the New Mexican 
Wasatch. Metatype, No. 4262 an incomplete skull, jaws and parts of skeleton from 
the Big Horn Wasatch. 
Type of P. intermedia, No. 2854 an upper jaw fragment with m?~ from the Big 
Horn Wasatch. 
Distinctive characters: Size-medium, lower jaw 360 mm. in total length. Skull 
and jaws elongate, canines massive, condyles heavy. Limbs long and moderately 
slender, feet digitigrade tetradactyl. Metacones of upper molars moderately reduced, 
m3 smaller than m! but variable in size. 
A skull, No. 15730, and a skeleton, No. 16154, from the Gray Bull beds 
are referred to this species. Both are fragmentary and incomplete but the 
bone well preserved and uncrushed, and they add considerable to what has 
hitherto been known of the morphology of Pachyena. A number of upper 
and lower jaws and jaw fragments are also referable. 
Pachyena intermedia of Wortman I am unable to separate specifically 
from P. ossifraga, although it is somewhat smaller than the skull and 
skeleton described by Cope (A. M. No. 4262) and the last molar slightly 
more reduced. It agrees more nearly with the skull and skeleton Nos. 
15730, 16154, which in turn agree closely with the type of P. ossifraga. No. 
4262 is a more robust individual, and the series of upper and lower jaws 
referred to the species show all kinds of intermediate conditions between 
these extremes. The hind limb referred to intermedia by Matthew in 1909 
is undoubtedly of a distinct species from ossifraga; but it belongs not to 
intermedia, but to the smaller species P. gracilis described above. 
The skull, No. 15730, as restored, is of very peculiar proportions. The 
mesocranial region is greatly elongate, the distance between m; and the 
postglenoid process exceeding the distance from canine to ms; _ the glenoid 
articulations are very large, project far downward, and are, for a carnivore, 
set far back. The anterior border of the orbit is above the posterior end of 
m’, The posterior nares open a little behind m3, are very narrow and 
constricted, the pterygoid plates set near together. The proportions of 
the skull resemble those of Harpagolestes rather than Mesonyx. In the Cope 
skull (No. 4262) the posterior part has been telescoped and crowded together 
by crushing so that these proportions are not brought out. The skull of 
H. macrocephalus as figured by Wortman appears to be of about the same 
proportions except that the face is shorter and. broader. 
The feet in No. 16154 are nearly as progressive as those of Synoplo- 
